Pollen are the tectum and the foot layer, which is just above the intine. The tectum and foot layer are separated by a region called the columella, which is composed of strengthening rods. The outer wall is constructed with a resistant biopolymer called sporopollenin. Pollen apertures are regions of the pollen wall that may involve exine thinning or a significant reduction in exine thickness. They allow shrinking and swelling of the grain caused by changes in moisture content. Elongated apertures or furrows in the pollen grain are called colpi (singular: colpus) or sulci (singular: sulcus). Apertures that are more circular are
Tylosis (botany) Tylosis (botany) A tylosis (plural: tyloses) is a bladder-like distension of a parenchyma cell into the lumen of adjacent vessels in wood. Less certainly, the term tylosis is in use to summarise the physiological process and the resulting occlusion in the xylem of woody plants as response to injury or as protection from decay in heartwood. It is a key process in wall one of the Compartmentalization Of Decay In Trees (CODIT) and other woody plants. Observed in section under a microscope, tyloses appear as balloon-like protrusions emanating from axial paratracheal parenchyma cells into xylem vessels through pits linking the