Pollen are the tectum and the foot layer, which is just above the intine. The tectum and foot layer are separated by a region called the columella, which is composed of strengthening rods. The outer wall is constructed with a resistant biopolymer called sporopollenin. Pollen apertures are regions of the pollen wall that may involve exine thinning or a significant reduction in exine thickness. They allow shrinking and swelling of the grain caused by changes in moisture content. Elongated apertures or furrows in the pollen grain are called colpi (singular: colpus) or sulci (singular: sulcus). Apertures that are more circular are
Cortex (botany) cylinder of the root through diffusion and may also be used for food storage in the form of starch. Cortex (botany) A cortex is the outermost layer of a stem or root in a plant. In botany, the cortex is the outermost layer of the stem or root of a plant, bounded on the outside by the epidermis and on the inside by the endodermis. In plants, it is composed mostly of differentiated cells, usually large thin-walled parenchyma cells of the ground tissue system. The outer cortical cells often acquire irregularly thickened cell walls, and are called collenchyma cells. Some