Pollination cones (female), but some species are monoecious and others dioecious. A pollen cone contains hundreds of microsporangia carried on (or borne on) reproductive structures called sporophylls. Spore mother cells in the microsporangia divide by meiosis to form haploid microspores that develop further by two mitotic divisions into immature male gametophytes (pollen grains). The four resulting cells consist of a large tube cell that forms the pollen tube, a generative cell that will produce two sperm by mitosis, and two prothallial cells that degenerate. These cells comprise a very reduced microgametophyte, that is contained within the resistant wall of the pollen
Tylosis (botany) then dies as its contents are disgorged into the empty space of the dry vessel and highly effective decay-inhibiting substances, notably tannin, are formed and absorbed by the adjacent vessel walls. Tylosis (botany) A tylosis (plural: tyloses) is a bladder-like distension of a parenchyma cell into the lumen of adjacent vessels in wood. Less certainly, the term tylosis is in use to summarise the physiological process and the resulting occlusion in the xylem of woody plants as response to injury or as protection from decay in heartwood. It is a key process in wall one of the Compartmentalization Of Decay