Pollen cells are surrounded by a thin delicate wall of unaltered cellulose called the endospore or intine, and a tough resistant outer cuticularized wall composed largely of sporopollenin called the exospore or exine. The exine often bears spines or warts, or is variously sculptured, and the character of the markings is often of value for identifying genus, species, or even cultivar or individual. The spines may be less than a micron in length (spinulus, plural spinuli) referred to as spinulose (scabrate), or longer than a micron (echina, echinae) referred to as echinate. Various terms also describe the sculpturing such as reticulate,
Tylosis (botany) then dies as its contents are disgorged into the empty space of the dry vessel and highly effective decay-inhibiting substances, notably tannin, are formed and absorbed by the adjacent vessel walls. Tylosis (botany) A tylosis (plural: tyloses) is a bladder-like distension of a parenchyma cell into the lumen of adjacent vessels in wood. Less certainly, the term tylosis is in use to summarise the physiological process and the resulting occlusion in the xylem of woody plants as response to injury or as protection from decay in heartwood. It is a key process in wall one of the Compartmentalization Of Decay