Pollen cells are surrounded by a thin delicate wall of unaltered cellulose called the endospore or intine, and a tough resistant outer cuticularized wall composed largely of sporopollenin called the exospore or exine. The exine often bears spines or warts, or is variously sculptured, and the character of the markings is often of value for identifying genus, species, or even cultivar or individual. The spines may be less than a micron in length (spinulus, plural spinuli) referred to as spinulose (scabrate), or longer than a micron (echina, echinae) referred to as echinate. Various terms also describe the sculpturing such as reticulate,
Hilum (biology) such a mark, and can be used as a distinguishing characteristic of a seed or spore. Hilum (biology) In botany, a hilum (pronounced ) is a scar or mark left on a seed coat by the former attachment to the ovary wall or to the funiculus (which in turn attaches to the ovary wall). On a bean seed, the hilum is called the "eye". For some species of fungus, the hilum is the microscopic indentation left on a spore when it separates from the sterigma of the basidium. A hilum can also be a nucleus of a starch grain; the