Pollen grains often have air bladders, or vesicles, called sacci. The sacci are not actually balloons, but are sponge-like, and increase the buoyancy of the pollen grain and help keep it aloft in the wind, as most gymnosperms are anemophilous. Pollen can be monosaccate, (containing one saccus) or bisaccate (containing two sacci). Modern pine, spruce, and yellowwood trees all produce saccate pollen. The transfer of pollen grains to the female reproductive structure (pistil in angiosperms) is called pollination. This transfer can be mediated by the wind, in which case the plant is described as anemophilous (literally wind-loving). Anemophilous plants typically produce
Medullary ray (botany) also known as vascular rays or pith rays. In this context, the term refers to radial sheets or ribbons extending vertically through the tree across and perpendicular to the growth rings. Also called pith rays or wood rays, these formations of primarily parenchyma cells allow the radial transport of sap and are essential in the process of tylosis. In quartersawn material, where the wood is cut into boards with the growth rings roughly perpendicular to the face of the board, the medullary rays often produce beautiful figure such as silver grain, medullary spots, pith flecks, etc. Medullary ray (botany) Medullary