Pollen cells are surrounded by a thin delicate wall of unaltered cellulose called the endospore or intine, and a tough resistant outer cuticularized wall composed largely of sporopollenin called the exospore or exine. The exine often bears spines or warts, or is variously sculptured, and the character of the markings is often of value for identifying genus, species, or even cultivar or individual. The spines may be less than a micron in length (spinulus, plural spinuli) referred to as spinulose (scabrate), or longer than a micron (echina, echinae) referred to as echinate. Various terms also describe the sculpturing such as reticulate,
Medullary ray (botany) also known as vascular rays or pith rays. In this context, the term refers to radial sheets or ribbons extending vertically through the tree across and perpendicular to the growth rings. Also called pith rays or wood rays, these formations of primarily parenchyma cells allow the radial transport of sap and are essential in the process of tylosis. In quartersawn material, where the wood is cut into boards with the growth rings roughly perpendicular to the face of the board, the medullary rays often produce beautiful figure such as silver grain, medullary spots, pith flecks, etc. Medullary ray (botany) Medullary