Pollen grains often have air bladders, or vesicles, called sacci. The sacci are not actually balloons, but are sponge-like, and increase the buoyancy of the pollen grain and help keep it aloft in the wind, as most gymnosperms are anemophilous. Pollen can be monosaccate, (containing one saccus) or bisaccate (containing two sacci). Modern pine, spruce, and yellowwood trees all produce saccate pollen. The transfer of pollen grains to the female reproductive structure (pistil in angiosperms) is called pollination. This transfer can be mediated by the wind, in which case the plant is described as anemophilous (literally wind-loving). Anemophilous plants typically produce
Cortex (botany) Cortex (botany) A cortex is the outermost layer of a stem or root in a plant. In botany, the cortex is the outermost layer of the stem or root of a plant, bounded on the outside by the epidermis and on the inside by the endodermis. In plants, it is composed mostly of differentiated cells, usually large thin-walled parenchyma cells of the ground tissue system. The outer cortical cells often acquire irregularly thickened cell walls, and are called collenchyma cells. Some of the outer cortical cells may contain chloroplasts. It is responsible for the transportation of materials into the central