Pollen called pores. Colpi, sulci and pores are major features in the identification of classes of pollen. Pollen may be referred to as inaperturate (apertures absent) or aperturate (apertures present). The aperture may have a lid (operculum), hence is described as operculate. However the term inaperturate covers a wide range of morphological types, such as functionally inaperturate (cryptoaperturate) and omniaperturate. Inaperaturate pollen grains often have thin walls, which facilitates pollen tube germination at any position. Terms such as uniaperturate and triaperturate refer to the number of apertures present (one and three respectively). The orientation of furrows (relative to the original tetrad
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