Pollen are the tectum and the foot layer, which is just above the intine. The tectum and foot layer are separated by a region called the columella, which is composed of strengthening rods. The outer wall is constructed with a resistant biopolymer called sporopollenin. Pollen apertures are regions of the pollen wall that may involve exine thinning or a significant reduction in exine thickness. They allow shrinking and swelling of the grain caused by changes in moisture content. Elongated apertures or furrows in the pollen grain are called colpi (singular: colpus) or sulci (singular: sulcus). Apertures that are more circular are
Decay theory that older memories (in long-term memory) are often more resistant to shocks or physical attacks on the brain. It is also thought that the passage of time alone cannot cause forgetting, and that decay theory must also take into account some processes that occur as more time passes. The term "decay theory" was first coined by Edward Thorndike in his book "The Psychology of Learning" in 1914. This simply states that if a person does not access and use the memory representation they have formed the memory trace will fade or decay over time. This theory was based on the