Pollen grains often have air bladders, or vesicles, called sacci. The sacci are not actually balloons, but are sponge-like, and increase the buoyancy of the pollen grain and help keep it aloft in the wind, as most gymnosperms are anemophilous. Pollen can be monosaccate, (containing one saccus) or bisaccate (containing two sacci). Modern pine, spruce, and yellowwood trees all produce saccate pollen. The transfer of pollen grains to the female reproductive structure (pistil in angiosperms) is called pollination. This transfer can be mediated by the wind, in which case the plant is described as anemophilous (literally wind-loving). Anemophilous plants typically produce
Tylosis (botany) then dies as its contents are disgorged into the empty space of the dry vessel and highly effective decay-inhibiting substances, notably tannin, are formed and absorbed by the adjacent vessel walls. Tylosis (botany) A tylosis (plural: tyloses) is a bladder-like distension of a parenchyma cell into the lumen of adjacent vessels in wood. Less certainly, the term tylosis is in use to summarise the physiological process and the resulting occlusion in the xylem of woody plants as response to injury or as protection from decay in heartwood. It is a key process in wall one of the Compartmentalization Of Decay