Vittarioideae worthy of subgeneric rank, while "Rheopteris" had not yet been discovered. "Haplopteris" he explicitly synonymized with "Vittaria" in 1914. Carl Christensen used the name "Vittarioideae" in Verdoorn's "Manual of Pteridology" in 1938, but did not include a description, leaving it nomenclaturally invalid. Ren-Chang Ching raised Vittariaceae to the rank of a family in 1940. The first well-sampled molecular phylogenetic study of the vittarioids was based on the chloroplast gene rbcL. In this study, it was found that the type species of "Monogramma" is embedded in "Haplopteris"; the segregation of "Vaginularia" from "Monogramma" was also supported, as members of "Vaginularia" formed
Evolutionary history of plants in place of flowers. Thus, sometime in history, the developmental program leading to formation of a leaf must have been altered to generate a flower. There probably also exists an overall robust framework within which the floral diversity has been generated. An example of that is a gene called "LEAFY (LFY)", which is involved in flower development in "Arabidopsis thaliana". The homologs of this gene are found in angiosperms as diverse as tomato, snapdragon, pea, maize and even gymnosperms. Expression of "Arabidopsis thaliana" LFY in distant plants like poplar and citrus also results in flower-production in these plants. The "LFY"