Evolutionary history of plants and radially symmetrical, is an integumented megasporangium surrounded by a cupule. The megasporangium bears an unopened distal extension protruding above the multilobed integument. It is suspected that the extension was involved in anemophilous pollination. "Runcaria" sheds new light on the sequence of character acquisition leading to the seed. "Runcaria" has all of the qualities of seed plants except for a solid seed coat and a system to guide the pollen to the ovule. The first spermatophytes (literally: "seed plants") – that is, the first plants to bear true seeds – are called pteridosperms: literally, "seed ferns", so called because their
Metamorphosis of Plants than fifty years earlier. Metamorphosis of Plants Versuch die Metamorphose der Pflanzen zu erklären, known in English as Metamorphosis of Plants, was published by German poet and philosopher Johann Wolfgang von Goethe in 1790. In this work, Goethe essentially discovered the (serially) homologous nature of leaf organs in plants, from cotyledons, to photosynthetic leaves, to the petals of a flower. Although Sir Richard Owen, the British vertebrate anatomist, is generally credited with first articulating a definition of the word "homology" (in 1843), it is clear that Goethe had already arrived at a sophisticated view of homology and transformation (within an