Vittarioideae worthy of subgeneric rank, while "Rheopteris" had not yet been discovered. "Haplopteris" he explicitly synonymized with "Vittaria" in 1914. Carl Christensen used the name "Vittarioideae" in Verdoorn's "Manual of Pteridology" in 1938, but did not include a description, leaving it nomenclaturally invalid. Ren-Chang Ching raised Vittariaceae to the rank of a family in 1940. The first well-sampled molecular phylogenetic study of the vittarioids was based on the chloroplast gene rbcL. In this study, it was found that the type species of "Monogramma" is embedded in "Haplopteris"; the segregation of "Vaginularia" from "Monogramma" was also supported, as members of "Vaginularia" formed
Metamorphosis of Plants than fifty years earlier. Metamorphosis of Plants Versuch die Metamorphose der Pflanzen zu erklären, known in English as Metamorphosis of Plants, was published by German poet and philosopher Johann Wolfgang von Goethe in 1790. In this work, Goethe essentially discovered the (serially) homologous nature of leaf organs in plants, from cotyledons, to photosynthetic leaves, to the petals of a flower. Although Sir Richard Owen, the British vertebrate anatomist, is generally credited with first articulating a definition of the word "homology" (in 1843), it is clear that Goethe had already arrived at a sophisticated view of homology and transformation (within an