Vittarioideae worthy of subgeneric rank, while "Rheopteris" had not yet been discovered. "Haplopteris" he explicitly synonymized with "Vittaria" in 1914. Carl Christensen used the name "Vittarioideae" in Verdoorn's "Manual of Pteridology" in 1938, but did not include a description, leaving it nomenclaturally invalid. Ren-Chang Ching raised Vittariaceae to the rank of a family in 1940. The first well-sampled molecular phylogenetic study of the vittarioids was based on the chloroplast gene rbcL. In this study, it was found that the type species of "Monogramma" is embedded in "Haplopteris"; the segregation of "Vaginularia" from "Monogramma" was also supported, as members of "Vaginularia" formed
Evolutionary history of plants variance in the population. Therefore, if there is only a small population of plants in an area with the ability to reproduce together, genetic drift may counteract the effects of selection putting the plant in a disadvantageous position to fungi which can evolve at a normal rate. The variance in both the host and pathogen population is a major determinant of evolutionary success compared to the other species. The greater the genetic variance, the faster the species can evolve to counteract the other organism’s avoidance or defensive mechanisms. Due to the process of pollination for plants, the effective population size