Akodon spegazzinii is found in northwestern Argentina, in the provinces of Salta, Catamarca, Tucumán, La Rioja, and Mendoza, at altitudes of . Although its main distribution is in the northern provinces of Salta, Tucumán, and Catamarca, there are also scattered records from the more southern provinces of La Rioja and Mendoza, where it is likely restricted to patches of wet habitat. "Akodon alterus" has been reported from Jujuy, but this record was likely based on misidentified specimens of "A. boliviensis", and records of "Akodon spegazzinii" from Jujuy were based on misidentified "A. sylvanus". "Akodon spegazzinii" is known from a paleontological site in
Ruben Mendoza (American football) coach and also coached the defensive line on the school's football team. four years as the Coordinator of Strength and Conditioning at the University of Tennessee before moving to Clemson University to serve as Assistant Director of Strength and Conditioning. Mendoza later spent four years as Director of Strength and Conditioning at the University of Mississippi before being named Strength and Conditioning Coordinator at the University of Notre Dame. He is now the head strength and conditioning coach at Wayne State University in Detroit. Ruben Mendoza (American football) Ruben Mendoza is a strength and conditioning coach and a former guard