Opuntia including antelopes, non-human primates, elephants, birds, and humans. When Carl Linnaeus published "Species Plantarum" in 1753 – the starting point for modern botanical nomenclature – he placed all the species of cactus known to him in one genus, "Cactus". In 1754, the Scottish botanist Philip Miller divided cacti into several genera, including "Opuntia". He distinguished the genus largely on the form of its flowers and fruits. There is considerable variation of taxonomy within "Opuntia" species, resulting in names being created for variants or subtypes within a species, and use of DNA sequencing to define and isolate various species. "Opuntia" hybridizes
Opuntia engelmannii (state) to the Tamaulipan matorral in Chihuahua (state), in Mexico. In the Sonoran Desert, terminal pads face predominantly east-west, so as to maximize the absorption of solar radiation during summer rains. Although found occasionally in the Mojave Desert, it tends to be replaced by "Opuntia basilaris", which does not need the summer rain. The overall form of "Opuntia engelmannii" is generally shrubby, with dense clumps up to high, usually with no apparent trunk. The pads are green (rarely blue-green), obovate to round, about 15–30 cm long and 12–20 cm wide. The glochids are yellow initially, then brown with age. Spines