Seabird in the Cretaceous, with a species called "Tytthostonyx glauconiticus", which seems allied to the Procellariiformes and Pelecaniformes. In the Paleogene both pterosaurs and marine reptiles became extinct, allowing seabirds to expand ecologically. These post-extinction seas were dominated by early Procellariidae, giant penguins and two extinct families, the Pelagornithidae and the Plotopteridae (a group of large seabirds that looked like the penguins). Modern genera began their wide radiation in the Miocene, although the genus "Puffinus" (which includes today's Manx shearwater and sooty shearwater) might date back to the Oligocene. The highest diversity of seabirds apparently existed during the Late Miocene and
Procellariidae with "Pseudobulweria" being phylogenetically closer to the "Puffinus" shearwaters than the "Pterodroma" gadfly petrels, and "Aphrodroma" (the Kerguelen petrel) possibly being closely related to the shearwaters or the fulmars. The prions, according to Nunn and Stanley, were among the larger shearwater group. The "Calonectris" shearwaters were placed close to the two "Puffinus" clades (closer to the Puffinus, or small, clade) and both were distant to the "Procellaria" shearwaters. The relationships between the genera and within the genera are still the subject of debate, with researchers lumping and splitting the species and genera within the family and arguing about the position