Puffin are closely related, together composing the subfamily Fraterculini. The scientific name "Fratercula" is derived from Latin and means "little brother", a reference to the black and white plumage, which resembles monastic robes. The English name "puffin" – puffed in the sense of swollen – was originally applied to the fatty, salted meat of young birds of the unrelated Manx shearwater ("Puffinus puffinus"), formerly known as the "Manks puffin". Puffin is an Anglo-Norman word (Middle English "pophyn" or "poffin") for the cured carcasses of nestling Manx shearwaters. The oldest alcid fossil is "Hydrotherikornis" from Oregon dating to the Late Eocene while
Dune shearwater which bred in lava fields. Extinction of the species occurred about 2000–3000 years ago, contemporaneously with the first human settlement of the islands by the Guanches, with the cause likely to be human predation. Dune shearwater The dune shearwater ("Puffinus holeae"), also known as the Canarian shearwater or Hole's shearwater, was a relatively large shearwater which bred in the Canary Islands archipelago of the North Atlantic Ocean. Fossils have also been found in the Figueira Brava cave archaeological site on the western coast of Portugal. The specific epithet honours Mrs Jean Hole who collected fossil material of the species on