Sun compass in animals internal clock to determine their heading, and that they had learnt the actual direction down to the sea on their particular beach. Lockley's experiments with Manx shearwaters ("Puffinus puffinus") showed that when released "under a clear sky" far from their nests (in Skokholm), the seabirds first oriented themselves and then "flew off in a direct line for Skokholm", making the journey rapidly. For example, one of the birds, released at Boston airport, arrived in Skokholm 12½ days later; Lockley calculated that if the birds flew for 12 hours per day, they must have travelled at 20 miles per hour, their
Procellariidae with "Pseudobulweria" being phylogenetically closer to the "Puffinus" shearwaters than the "Pterodroma" gadfly petrels, and "Aphrodroma" (the Kerguelen petrel) possibly being closely related to the shearwaters or the fulmars. The prions, according to Nunn and Stanley, were among the larger shearwater group. The "Calonectris" shearwaters were placed close to the two "Puffinus" clades (closer to the Puffinus, or small, clade) and both were distant to the "Procellaria" shearwaters. The relationships between the genera and within the genera are still the subject of debate, with researchers lumping and splitting the species and genera within the family and arguing about the position