Seabird in the Cretaceous, with a species called "Tytthostonyx glauconiticus", which seems allied to the Procellariiformes and Pelecaniformes. In the Paleogene both pterosaurs and marine reptiles became extinct, allowing seabirds to expand ecologically. These post-extinction seas were dominated by early Procellariidae, giant penguins and two extinct families, the Pelagornithidae and the Plotopteridae (a group of large seabirds that looked like the penguins). Modern genera began their wide radiation in the Miocene, although the genus "Puffinus" (which includes today's Manx shearwater and sooty shearwater) might date back to the Oligocene. The highest diversity of seabirds apparently existed during the Late Miocene and
Persian shearwater off Oman and Socotra, Yemen, while "P. p. temptator" has been estimated to number fewer than 500 pairs on Moheli Island in the Comoros (reviewed by Brooke 2004). It is thought likely that more breeding colonies lie undiscovered elsewhere in the northern Indian Ocean (Onley and Scofield 2007), thus the total population probably includes more than 10,000 mature individuals. The population trend is believed to be downwards. Persian shearwater The Persian shearwater ("Puffinus persicus") is a seabird in the family Procellariidae formerly lumped in with Audubon's shearwater ("Puffinus lherminieri"). There are two listed subspecies of the Persian shearwater: After breeding,