Audubon's shearwater ill-resolved clade also including such species as the little shearwater, the Manx shearwater ("P. puffinus") or the black-vented shearwater ("P. opisthomelas"). The little-known Heinroth's shearwater ("P. heinrothi") is sometimes considered a subspecies of either Audubon's or the little shearwater. Though it is likely to be another member of that close-knit group, its actual relationships remain uncertain due to lack of specimens. Audubon's shearwater itself has around 10 subspecies. Several have at one time or another been suggested to constitute separate species. For example, the Galápagos Islands population has turned out to be a very distinct species, the Galápagos shearwater ("P.
Dune shearwater which bred in lava fields. Extinction of the species occurred about 2000–3000 years ago, contemporaneously with the first human settlement of the islands by the Guanches, with the cause likely to be human predation. Dune shearwater The dune shearwater ("Puffinus holeae"), also known as the Canarian shearwater or Hole's shearwater, was a relatively large shearwater which bred in the Canary Islands archipelago of the North Atlantic Ocean. Fossils have also been found in the Figueira Brava cave archaeological site on the western coast of Portugal. The specific epithet honours Mrs Jean Hole who collected fossil material of the species on