The Liver Birds and Series 1 starred Pauline Collins as Dawn and Polly James as Beryl Hennessey. In Series 2, Nerys Hughes debuted as Sandra Hutchinson, replacing Dawn for the rest of the programme's run. The Beryl-and-Sandra pairing is generally regarded as the programme's best period. Beryl was the more 'common' one, while Sandra was soft-spoken and more refined, due to the influence of her snobbish, overbearing mother (Mollie Sugden). Carla Lane drew on her own mother for the character – "Mrs Hutchinson, I think she was my mother. I'm sure she was my mother". Beryl's 'common' mother (the Hennesseys live in Bootle,
The Liver Birds a working class district north of the city) was played by Sheila Fay. Future "Emmerdale" actor Elizabeth Estensen as Carol Boswell replaced Beryl from Series 5 onwards. The title comes from the name given to two sculpted birds perched on top of the Royal Liver Building at Pier Head in the city of Liverpool. Michael Mills, who commissioned the series, came up with the title, although it was initially disliked by Carla Lane. The Scaffold—a pop group including Mike McCartney (brother of former Beatle Paul McCartney), poet Roger McGough and John Gorman of "Tiswas" fame—sang the title song. The title