Wookey Hole Caves then passes through major cave systems such as Swildon's Hole, Eastwater Cavern and St Cuthbert's Swallet, around Priddy, but 95% is water that has percolated directly into the limestone. The southern slopes of the Mendip Hills largely follow the flanks of an anticline, a fold in the rock that is convex upwards and has its oldest beds at its core. On the Mendips the crest of the anticline is truncated by erosion, forming a plateau. The rock strata here dip 10–15 degrees to the southwest. The outer slopes are mainly of Carboniferous Limestone, with Devonian age Old Red Sandstone exposed
Wookey Hole Caves plateau before sinking at the limestone boundary into cave systems such as Swildon's Hole, Eastwater Cavern and St Cuthbert's Swallet; the rest is rain that percolates directly through the limestone. The temperature in the caves is a constant . The caves have been used by humans for around 45,000 years, demonstrated by the discovery of tools from the Palaeolithic period, along with fossilised animal remains. Evidence of Stone and Iron Age occupation continued into Roman Britain. A corn-grinding mill operated on the resurgent waters of the River Axe as early as the Domesday survey of 1086. The waters of the