Elsa the lioness the Meru National Park. She died as local sentiment had begun to turn against Elsa and her cubs, forcing the Adamsons to consider moving them. Elsa’s death made her cubs much more averse to human contact, even with the Adamsons themselves, complicating what would be their eventual capture and release in the Serengeti. The fate of the cubs upon their release was uncertain, though George Adamson was able to find Little Elsa, healthy and in the company of two other unrelated lions, during 19 months of subsequent searching. This was the last time that the Adamsons saw any of Elsa's
Mara the Lioness earlier when they had attempted to release her back into the wild. Eventually only Ugas was released into the Meru Game Reserve. It was decided that Mara was too tame and she and the other lionesses should be relocated to Whipsnade Park. Just prior to her departure Mara gave birth to male and female twin cubs, sired by Ugas during the filming of "Born Free". In 1966, Irene Grindlay wrote a book titled "Velvet Paws" which detailed the life of Mara up to and including her appearance in the movie "Born Free". ""We have no reason to reproach ourselves. We