Malus sylvestris to most European countries. It occurs in a scattered distribution pattern as single individuals or in small groups. In the past "M. sylvestris" was thought to be the most important ancestor of the cultivated apple ("M. pumila"), which has since been shown to have been originally derived from the central Asian species "M. sieversii". However another recent DNA analysis confirms that "M. sylvestris" has contributed significantly to the genome. The study found that secondary introgression from other species of the genus "Malus" has greatly shaped the genome of "M. pumila", with "M. sylvestris" being the largest secondary contributor. It also
Karl Malus U.S. Agent then destroyed Malus' equipment and records. Malus has since worked for a variety of criminal organizations, including the Corporation, and the Maggia. He even worked with the Avengers and the Thunderbolts in their efforts to defeat Count Nefaria in exchange for a reduced sentence. During the "Dark Reign" storyline, Karl Malus was researched by Quasimodo as part of its work for Norman Osborn. Karl Malus was recruited by the Wizard to become a new member of the Frightful Four, only to find himself being forcibly made the new host of the Carnage Symbiote. He was eaten by Carnage