Evolution of the cochlea and the elongated cochlea allows for better sensitivity for higher frequencies. As in all lepidosaurs and archosaurs, the single-ossicle (columellar) middle ear transmits sound to the footplate of the columella, which sends a pressure wave through the inner ear. In snakes, the basilar papilla is roughly 1mm long and only responds to frequencies below about 1 kHz. In contrast, lizards tend to have two areas of hair cells, one responding below and the other above 1 kHz. The upper frequency limit in most lizards is roughly 5–8 kHz. The longest lizard papillae are about 2mm long and contain 2000 hair
Evolution of the cochlea cells and their afferent innervating fibers can be very sharply tuned to frequency. In birds and crocodilians, the similarity of the structure of the basilar papilla betrays their close evolutionary relationship. The basilar papilla is up to about 10mm long and contains up to 16500 hair cells. While most birds have an upper hearing limit of only about 6 kHz, the barn owl can hear up to 12 kHz and thus close to the human upper limit. Egg-laying mammals, the monotremes (spiny anteater and platypus), do not have a spiral cochlea, but one shaped more like a banana, up to