Canary Islands be many years before the island was fully conquered. The natives of La Gomera, and of Gran Canaria, Tenerife, and La Palma, resisted the Castilian invaders for almost a century. In 1448 Maciot de Béthencourt sold the lordship of Lanzarote to Portugal's Prince Henry the Navigator, an action that was accepted by neither the natives nor the Castilians. Despite Pope Nicholas V ruling that the Canary Islands were under Portuguese control, the crisis swelled to a revolt which lasted until 1459 with the final expulsion of the Portuguese. In 1479, Portugal and Castile signed the Treaty of Alcáçovas, which settled
Gallotia 2010). Finally, the clade that colonized the younger western islands was likely to have originated from Lanzarote/Fuerteventura. This western island clade diverged into two groups, all of which colonized Tenerife, La Palma, La Gomera and El Hierro, leading to 1) a medium-bodied (e.g., "G. caesaris" from El Hierro) and 2) a large bodied "giant" species (e.g., "G. simonyi" from El Hierro) on each of these islands (note that "G. intermedia" from Tenerife belongs to the "giant" group, but present-day individuals are not that large). The giant species now exist, at best, in small relict populations, while "G. auritae" may be