Evolution of the cochlea about 7 mm long. Like in lepidosaurs and archosaurs, it contains a lagena, a vestibular sensory epithelium, at its tip. Only in therian mammals (marsupials and placentals) is the cochlea truly coiled 1.5 to 3.5 times. Whereas in monotremes there are many rows of both inner and outer hair cells in the organ of Corti, in therian (marsupial and placental) mammals the number of inner hair-cell rows is one, and there are generally only three rows of outer hair cells. Amphibians have unique inner ear structures. There are two sensory papillae involved in hearing, the basilar (higher frequency) and amphibian
Evolution of the cochlea (lower frequency) papillae, but it is uncertain whether either is homologous to the hearing organs of lepidosaurs, archosaurs and mammals and we have no idea when they arose. Fish have no dedicated auditory epithelium, but use various vestibular sensory organs that respond to sound. In most teleost fishes it is the saccular macula that responds to sound. In some, such as goldfishes, there is also a special bony connection to the gas bladder that increases sensitivity allowing hearing up to about 4 kHz. The size of cochlea has been measured throughout its evolution based on the fossil record. In one