Evolution of the cochlea and the elongated cochlea allows for better sensitivity for higher frequencies. As in all lepidosaurs and archosaurs, the single-ossicle (columellar) middle ear transmits sound to the footplate of the columella, which sends a pressure wave through the inner ear. In snakes, the basilar papilla is roughly 1mm long and only responds to frequencies below about 1 kHz. In contrast, lizards tend to have two areas of hair cells, one responding below and the other above 1 kHz. The upper frequency limit in most lizards is roughly 5–8 kHz. The longest lizard papillae are about 2mm long and contain 2000 hair
Evolution of the cochlea apex of the cochlea faces more inferiorly in the hominins than the latter two groups. Finally, the cochlea of European middle Pleistocene hominins faces more inferiorly than Neanderthals, modern humans, and Homo erectus. Human beings, along with Apes, are the only mammals that do not have high frequency (>32 kHz) hearing. Humans have long cochleae, but the space devoted to each frequency range is quite large (2.5mm per octave), resulting in a comparatively reduced upper frequency limit. The human cochlea has approximately 2.5 turns around the modiolus (the axis). Humans, like many mammals and birds, are able to perceive auditory