Evolution of the cochlea signals that displace the eardrum by a mere picometre. Because of its prominence and preserved state in the fossil record, until recently, the ear had been used to determine phylogeny. The ear itself contains different portions, including the outer ear, the middle ear, and the inner ear and all of these show evolutionary changes that are often unique to each lineage [14]. It was the independent evolution of a tympanic middle ear in the Triassic era that produced strong selection pressures towards improved hearing organs in the separate lineages of land vertebrates. The cochlea is the tri-chambered auditory detection portion
Evolution of the cochlea mammals. Prestins are located in the outer hair cells of mammalian cochlea and are considered motor proteins. They are found in the hair cells of all vertebrates, including fish, but are thought to have initially been membrane transporter molecules. A high concentration of prestins are found only in the lateral membranes of therian outer hair cells (there is uncertainty with regard to concentrations in monotremes). This high concentration is not found in inner hair cells, and is also lacking in all hair cell types of non-mammals. Prestin also has a role in motility, which evolved a greater importance in the