Evolution of the cochlea and the elongated cochlea allows for better sensitivity for higher frequencies. As in all lepidosaurs and archosaurs, the single-ossicle (columellar) middle ear transmits sound to the footplate of the columella, which sends a pressure wave through the inner ear. In snakes, the basilar papilla is roughly 1mm long and only responds to frequencies below about 1 kHz. In contrast, lizards tend to have two areas of hair cells, one responding below and the other above 1 kHz. The upper frequency limit in most lizards is roughly 5–8 kHz. The longest lizard papillae are about 2mm long and contain 2000 hair
Evolution of the cochlea at the same time as the water-to-land transition of vertebrates, approximately 380 million years ago. The actual coiling or spiral nature of the cochlea occurred to save space inside the skull. The longer the cochlea, the higher is the potential resolution of sound frequencies given the same hearing range. The oldest of the truly coiled mammalian cochleae were approximately 4 mm in length. The earliest evidence available for primates depicts a short cochlea with prominent laminae, suggesting that they had good high-frequency sensitivity as opposed to low-frequency sensitivity. After this, over a period of around 60 million years, evidence suggests