Evolution of the cochlea and the elongated cochlea allows for better sensitivity for higher frequencies. As in all lepidosaurs and archosaurs, the single-ossicle (columellar) middle ear transmits sound to the footplate of the columella, which sends a pressure wave through the inner ear. In snakes, the basilar papilla is roughly 1mm long and only responds to frequencies below about 1 kHz. In contrast, lizards tend to have two areas of hair cells, one responding below and the other above 1 kHz. The upper frequency limit in most lizards is roughly 5–8 kHz. The longest lizard papillae are about 2mm long and contain 2000 hair
Stirrup remained in use until the late Heian period when a new stirrup was developed. The "fukuro abumi" or "musashi abumi" had a base that extended the full length of the rider's foot and the right and left sides of the toe cup were removed. The open sides were designed to prevent the rider from catching a foot in the stirrup and being dragged. The military version of this open-sided stirrup ("shitanaga abumi") was in use by the middle Heian period. It was thinner, had a deeper toe pocket and an even longer and flatter foot shelf. This stirrup stayed in