Origin of the domestic dog or the result of natural introgression as the dog expanded across Eurasia. In 2018, a study found a small amount of dog ancestry in 62% of Eurasian wolf specimens looked at, that hybridization had occurred across a wide number of timescales and not just recently, however in contrast there was almost no admixture detected in the North American specimens. There was introgression of the male dog into the wolf, but also one hybrid detected which was the result of a male wolf crossed with a female dog. Wolves have maintained their phenotype differences from the dog, which indicates low-frequency hybridization.
Dog anatomy study of dog skull shape compared to extant carnivorans proposed that "The greatest shape distances between dog breeds clearly surpass the maximum divergence between species in the Carnivora. Moreover, domestic dogs occupy a range of novel shapes outside the domain of wild carnivorans." The domestic dog compared to the wolf shows the greatest variation in the size and shape of the skull (Evans 1979) that range from 7 to 28 cm in length (McGreevy 2004). Wolves are dolichocephalic (long skulled) but not as extreme as some breeds of dogs such as greyhounds and Russian wolfhounds (McGreevy 2004). Canine brachycephaly (short-skulledness)