Rust (fungus) only uredinia and telia. Spermagonia may be absent from each type but especially the microcyclic life cycle. In macrocyclic and demicyclic life cycles, the rust may be either host alternating (heteroecious), i.e., the aecial state is on one kind of plant but the telial state on a different and unrelated plant, or non-host alternating (autoecious), i.e., the aecial and telial states on the same kind of plant. Heteroecious rust fungi require two unrelated hosts to complete their life cycle, with the primary host being infected by aeciospores and the alternate host being infected with basidiospores. This can be contrasted with
Stem rust the end of the growing season black telia are produced. For this reason stem rust is also known as 'black rust'. The telia are firmly attached to the plant tissue. The site of infection is a visible symptom of the disease. Pycnia appear on barberry plants in the spring, usually in the upper leaf surfaces. They are often in small clusters and exude pycniospores in a sticky honeydew. Five to ten days later, cup-shaped structures filled with orange-yellow, powdery aeciospores break through the lower leaf surface. The aecial cups are yellow and sometimes elongate to extend up to 5 mm