Plant cell parenchyma cells, that covers the external surfaces of leaves, stems and roots. Several cell types may be present in the epidermis. Notable among these are the stomatal guard cells that control the rate of gas exchange between the plant and the atmosphere, glandular and clothing hairs or trichomes, and the root hairs of primary roots. In the shoot epidermis of most plants, only the guard cells have chloroplasts. Chloroplasts contain the green pigment chlorophyll which is needed for photosynthesis. The epidermal cells of aerial organs arise from the superficial layer of cells known as the "tunica" (L1 and L2 layers)
Granule cell fibers form thousands of excitatory Granule-cell-Purkinje-cell synapses onto the dendrites of Purkinje cells. This connection is excitatory as glutamate is released. The parallel fibers and ascending axon synapses from the same granule cell fire in synchrony which results in excitatory signals. In the cerebellar cortex there are a variety of inhibitory neurons (interneurons). The only excitatory neurons present in the cerebellar cortex are granule cells. Plasticity of the synapse between a parallel fiber and a Purkinje cell is believed to be important for motor learning. The function of cerebellar circuits is entirely dependent on processes carried out by the granular