Albertosaurus older "Gorgosaurus libratus" (sometimes called "Albertosaurus libratus"; see below). These two species are the only described albertosaurines; other undescribed species may exist. Thomas Holtz found "Appalachiosaurus" to be an albertosaurine in 2004, but his more recent unpublished work locates it just outside Tyrannosauridae, in agreement with other authors. The other major subfamily of tyrannosaurids is the Tyrannosaurinae, which includes "Daspletosaurus", "Tarbosaurus" and "Tyrannosaurus". Compared with these robust tyrannosaurines, albertosaurines had slender builds, with proportionately smaller skulls and longer bones of the lower leg (tibia) and feet (metatarsals and phalanges). Below is the cladogram of the Tyrannosauridae based on the phylogenetic
Dorsal interossei of the hand arising by two heads from the adjacent sides of the metacarpal bones, but more extensively from the metacarpal bone of the finger into which the muscle is inserted. They are inserted into the bases of the proximal phalanges and into the extensor expansion of the corresponding extensor digitorum tendon. The middle digit has two dorsal interossei insert onto it while the first digit (thumb) and the fifth digit (little finger) have none. Each finger is provided with two interossei (palmar or dorsal), with the exception of the little finger, in which the abductor digiti minimi muscle takes the place of