Albertosaurus older "Gorgosaurus libratus" (sometimes called "Albertosaurus libratus"; see below). These two species are the only described albertosaurines; other undescribed species may exist. Thomas Holtz found "Appalachiosaurus" to be an albertosaurine in 2004, but his more recent unpublished work locates it just outside Tyrannosauridae, in agreement with other authors. The other major subfamily of tyrannosaurids is the Tyrannosaurinae, which includes "Daspletosaurus", "Tarbosaurus" and "Tyrannosaurus". Compared with these robust tyrannosaurines, albertosaurines had slender builds, with proportionately smaller skulls and longer bones of the lower leg (tibia) and feet (metatarsals and phalanges). Below is the cladogram of the Tyrannosauridae based on the phylogenetic
Phalanx bone terminal phalanges and fused metacarpal bones able to absorb the stress from running. The sloth spends its life hanging upside-down from branches, and has highly specialized third and fourth digits for the purpose. They have short and squat proximal phalanges with much longer terminal phalanges. They have vestigial second and fifth metacarpals, and their palm extends to the distal interphalangeal joints. The arboreal specialization of these terminal phalanges makes it impossible for the sloth to walk on the ground where the animal has to drag its body with its claws. Phalanx bone The phalanges (singular: "phalanx" ) are digital bones