Origin of the domestic dog to the domestic dog and modern wolf lineages. The gray wolf is thought to be ancestral to the domestic dog, however its relationship to "C. variabilis", and the genetic contribution of "C. variabilis" to the dog, is the subject of debate. The Zhokhov Island (8,700 YBP) and Aachim (1,700 YBP) canid haplotypes fell within the domestic dog clade, cluster with S805, and also share their haplotypes with – or are one mutation away from – the Tibetan wolf ("C. l. chanco") and the recently-extinct Japanese wolf ("C. l. hodophilax"). This may indicate that these canids retained the genetic signature of
Origin of the domestic dog Bonn–Oberkassel dog buried beside humans 14,200 years ago to be the first undisputed dog, with disputed remains occurring 36,000 years ago. These dates imply that the earliest dogs arose in the time of human hunter-gatherers and not agriculturists. The dog was the first species to be domesticated. Where the genetic divergence of dog and wolf took place remains controversial, with the most plausible proposals spanning Western Europe, Central Asia and East Asia. This has been made more complicated by the most recent proposal that fits the available evidence, which is that an initial wolf population split into East and West