Digestive system of gastropods species that have undergone de-torsion, however, the oesophagus may open into the anterior of the stomach, which is therefore reversed from the usual gastropod arrangement. In "Tarebia granifera" the brood pouch is above the oesophagus. There is available an extensive rostrum on the anterior part of the oesophagus in all carnivorous gastropods. Some basal gastropod clades have oesophageal gland. In most species, the stomach itself is a relatively simple sac, and is the main site of digestion. In many herbivores, however, the hind part of the oesophagus is enlarged to form a crop, which, in terrestrial pulmonates, may even replace
Development of the digestive system from four components: the septum transversum (central tendon), the pleuroperitoneal membranes, the dorsal mesentery of the esophagus, and muscular components from somites at cervical levels three to five (C3–5) of the body wall. Since the septum transversum is located initially opposite cervical segments of three to five, and since muscle cells for the diaphragm originate from somites at these segments, the phrenic nerve, which innervates the diaphragm, also arises from these segments of the spinal cord (C3, 4, and 5). The thoracic cavity is divided into the pericardial cavity and two pleural cavities for the lungs by the pleuropericardial membranes.