Southern Ocean Southern Hemisphere, with the greatest concentration located on and around Antarctica. Four of the 18 penguin species live and breed on the mainland and its close offshore islands. Another four species live on the subantarctic islands. Emperor penguins have four overlapping layers of feathers, keeping them warm. They are the only Antarctic animal to breed during the winter. There are relatively few fish species in few families in the Southern Ocean. The most species-rich family are the snailfish (Liparidae), followed by the cod icefish (Nototheniidae) and eelpout (Zoarcidae). Together the snailfish, eelpouts and notothenioids (which includes cod icefish and several
Ratite as a whole probably had an origin in the northern hemisphere. Early Cenozoic northern hemisphere paleognaths such as "Lithornis", "Pseudocrypturus", "Paracathartes" and "Palaeotis" appear to be the most basal members of the clade. The various ratite lineages were probably descended from flying ancestors that colonised the south in multiple waves, probably initially in South America, and then evolved gigantism. Loss of flight allows birds to eliminate the energetic cost of maintaining flight-enabling pectoral muscle mass. The basal metabolic rate of volant species is much higher than that of flightless terrestrial birds. But energetic efficiency can only help explain the loss