Plant cell parenchyma cells, that covers the external surfaces of leaves, stems and roots. Several cell types may be present in the epidermis. Notable among these are the stomatal guard cells that control the rate of gas exchange between the plant and the atmosphere, glandular and clothing hairs or trichomes, and the root hairs of primary roots. In the shoot epidermis of most plants, only the guard cells have chloroplasts. Chloroplasts contain the green pigment chlorophyll which is needed for photosynthesis. The epidermal cells of aerial organs arise from the superficial layer of cells known as the "tunica" (L1 and L2 layers)
Evolution of photosynthesis C photosynthesis is the oldest and most common form. C3 is a plant that uses the calvin cycle for the initial steps that incorporate CO2 into organic material. C4 is a plant that prefaces the calvin cycle with reactions that incorporate CO2 into four-carbon compounds. CAM is a plant that uses crassulacean acid metabolism, an adaptation for photosynthesis in arid conditions. C4 and CAM Plants have special adaptations that save water. The biochemical capacity to use water as the source for electrons in photosynthesis evolved in a common ancestor of extant cyanobacteria. The geological record indicates that this transforming event