Palisade cell Palisade cell Palisade cells are plant cells located in leaves, right below the epidermis and cuticle. They are vertically elongated, a different shape from the spongy mesophyll cells beneath them in the leaf. Their chloroplasts absorb a major portion of the light energy used by the leaf. Palisade cells occur in dicotyledonous plants, and also in the net-veined monocots, the Araceae and Dioscoreaceae. Palisade cells contain the largest number of chloroplasts per cell, which makes them the primary site of photosynthesis in the leaves of those plants that contain them, converting the energy in light to the chemical energy of
Photosynthesis as ribulose bisphosphate (RuBP). Using the ATP and NADPH produced by the light-dependent reactions, the resulting compounds are then reduced and removed to form further carbohydrates, such as glucose. The first photosynthetic organisms probably evolved early in the evolutionary history of life and most likely used reducing agents such as hydrogen or hydrogen sulfide, rather than water, as sources of electrons. Cyanobacteria appeared later; the excess oxygen they produced contributed directly to the oxygenation of the Earth, which rendered the evolution of complex life possible. Today, the average rate of energy capture by photosynthesis globally is approximately 130 terawatts, which