Origin of the domestic dog to the domestic dog and modern wolf lineages. The gray wolf is thought to be ancestral to the domestic dog, however its relationship to "C. variabilis", and the genetic contribution of "C. variabilis" to the dog, is the subject of debate. The Zhokhov Island (8,700 YBP) and Aachim (1,700 YBP) canid haplotypes fell within the domestic dog clade, cluster with S805, and also share their haplotypes with – or are one mutation away from – the Tibetan wolf ("C. l. chanco") and the recently-extinct Japanese wolf ("C. l. hodophilax"). This may indicate that these canids retained the genetic signature of
Origin of the domestic dog have diverged from their ancestor at a later time. Immediately after divergence, the dog population outnumbered the wolf population, and later the dog population underwent a population reduction to be much lower than that of the wolf. In 2017, a study compared the nuclear genome sequences of three Neolithic dog specimens from Germany and Ireland with sequences from over 5,000 dogs and wolves. The study found that modern European dogs descended from their Neolithic ancestors with no evidence of a population turnover found by an earlier study. The study found that a single dog-wolf divergence occurred 36,900-41,500 YBP, followed by