Southern Ocean – fairly uniform temperatures of the deep ocean at the poles and the equator which differ by no more than 5 °C, and the major current systems or marine conveyor belt which transport egg and larva stages. The rocky shores of mainland Antarctica and its offshore islands provide nesting space for over 100 million birds every spring. These nesters include species of albatrosses, petrels, skuas, gulls and terns. The insectivorous South Georgia pipit is endemic to South Georgia and some smaller surrounding islands. Freshwater ducks inhabit South Georgia and the Kerguelen Islands. The flightless penguins are all located in the
Ratite as a whole probably had an origin in the northern hemisphere. Early Cenozoic northern hemisphere paleognaths such as "Lithornis", "Pseudocrypturus", "Paracathartes" and "Palaeotis" appear to be the most basal members of the clade. The various ratite lineages were probably descended from flying ancestors that colonised the south in multiple waves, probably initially in South America, and then evolved gigantism. Loss of flight allows birds to eliminate the energetic cost of maintaining flight-enabling pectoral muscle mass. The basal metabolic rate of volant species is much higher than that of flightless terrestrial birds. But energetic efficiency can only help explain the loss