Origin of the domestic dog or the result of natural introgression as the dog expanded across Eurasia. In 2018, a study found a small amount of dog ancestry in 62% of Eurasian wolf specimens looked at, that hybridization had occurred across a wide number of timescales and not just recently, however in contrast there was almost no admixture detected in the North American specimens. There was introgression of the male dog into the wolf, but also one hybrid detected which was the result of a male wolf crossed with a female dog. Wolves have maintained their phenotype differences from the dog, which indicates low-frequency hybridization.
Origin of the domestic dog is also such a difference between the domestication traits that researchers believe to have been essential at the early stages of domestication, and the improvement traits that have appeared since the split between wild and domestic populations. Domestication traits are generally fixed within all domesticates, and were selected during the initial episode of domestication of that animal or plant, whereas improvement traits are present only in a proportion of domesticates, though they may be fixed in individual breeds or regional populations. The dog was the first domesticant and was established across Eurasia before the end of the Late Pleistocene era,