Marine life appeared in the form of fish, which live exclusively in water. Some of these evolved into amphibians which spend portions of their lives in water and portions on land. Other fish evolved into land mammals and subsequently returned to the ocean as seals, dolphins or whales. Plant forms such as kelp and algae grow in the water and are the basis for some underwater ecosystems. Plankton, and particularly phytoplankton, are key primary producers forming the general foundation of the ocean food chain. Marine invertebrates exhibit a wide range of modifications to survive in poorly oxygenated waters including breathing tubes (see
Microbial mat the causes, cyanobacteria are the most self-sufficient of the mat organisms and were well-adapted to strike out on their own both as floating mats and as the first of the phytoplankton, which forms the basis of most marine food chains. The time at which eukaryotes first appeared is still uncertain: there is reasonable evidence that fossils dated between and represent eukaryotes, but the presence of steranes in Australian shales may indicate that eukaryotes were present . There is still debate about the origins of eukaryotes, and many of the theories focus on the idea that a bacterium first became an