Origin of the domestic dog to the domestic dog and modern wolf lineages. The gray wolf is thought to be ancestral to the domestic dog, however its relationship to "C. variabilis", and the genetic contribution of "C. variabilis" to the dog, is the subject of debate. The Zhokhov Island (8,700 YBP) and Aachim (1,700 YBP) canid haplotypes fell within the domestic dog clade, cluster with S805, and also share their haplotypes with – or are one mutation away from – the Tibetan wolf ("C. l. chanco") and the recently-extinct Japanese wolf ("C. l. hodophilax"). This may indicate that these canids retained the genetic signature of
Origin of the domestic dog could not be found in other dogs nor wolves today. As the taxonomic classification of the "proto-dog" Paleolithic dogs as being either dogs or wolves remains controversial, they were excluded from the study. The phylogenetic tree generated from mDNA sequences found a deep division between the Sarloos wolfdog and all other dogs, indicating that breed's recent deriving from the German Shepherd and captive gray wolves. The next largest division was between Eastern Asian dogs and Western Eurasian (Europe and the Middle East) dogs that had occurred between 14,000-6,400 YBP, with the Newgrange dog clustering with the Western Eurasian dogs. The