Southern Ocean Southern Hemisphere, with the greatest concentration located on and around Antarctica. Four of the 18 penguin species live and breed on the mainland and its close offshore islands. Another four species live on the subantarctic islands. Emperor penguins have four overlapping layers of feathers, keeping them warm. They are the only Antarctic animal to breed during the winter. There are relatively few fish species in few families in the Southern Ocean. The most species-rich family are the snailfish (Liparidae), followed by the cod icefish (Nototheniidae) and eelpout (Zoarcidae). Together the snailfish, eelpouts and notothenioids (which includes cod icefish and several
Palaeognathae flying birds that were common in the Cenozoic of the northern hemisphere, were also paleognaths. He argues that the lithornithiform bird "Paleotis", known from fossils in Denmark (northern hemisphere), shared unique anatomical features of the skull that make it a member of the same order as the ostriches. He also argued that the kiwis should not have reached New Zealand, which moved away from the mainland in the Early Cretaceous, if their ancestor was flightless; this claim at least has been vindicated by the discovery of the possibly volant "Proapteryx". He therefore deduced that lithornithiform ancestors could have reached the