Plant cell parenchyma cells, that covers the external surfaces of leaves, stems and roots. Several cell types may be present in the epidermis. Notable among these are the stomatal guard cells that control the rate of gas exchange between the plant and the atmosphere, glandular and clothing hairs or trichomes, and the root hairs of primary roots. In the shoot epidermis of most plants, only the guard cells have chloroplasts. Chloroplasts contain the green pigment chlorophyll which is needed for photosynthesis. The epidermal cells of aerial organs arise from the superficial layer of cells known as the "tunica" (L1 and L2 layers)
Fractionation of carbon isotopes in oxygenic photosynthesis However, growth rate alone does not account for observed fractionation. The flux of CO into and out of a cell is roughly proportional to the cell surface area, and the cell carbon biomass varies as a function of cell volume. Phytoplankton geometry that maximizes surface area to volume should have larger isotopic fractionation from photosynthesis. The biochemical characteristics of phytoplankton are similar to C3 plants, whereas the gas exchange characteristics more closely resemble the C4 strategy. More specifically, phytoplankton improve the efficiency of their primary carbon-fixing enzyme, RuBisCO, with carbon concentrating mechanisms (CCM), just as C4 plants accumulate CO in