Flightless bird that require a significant amount of overall metabolic energy, ratites decrease their basal metabolic rate and conserve energy. A study looking at the basal rates of birds found a significant correlation between low basal rate and pectoral muscle mass in kiwis. On the contrary, flightless penguins exude an intermediate basal rate. This is likely because penguins have well-developed pectoral muscles for hunting and diving in the water. For ground feeding birds, a cursorial lifestyle is more economical and allows for easier access to dietary requirements. Flying birds have different wing and feather structures that make flying easier, while flightless birds'
Flightless bird free to increase in abundance until the population was limited by food and territory. A study looking at energy conservation and the evolution of flightlessness hypothesized intraspecific competition selected for a reduced individual energy expenditure, which is achieved by the loss of flight. Some flightless varieties of island birds are closely related to flying varieties, implying flight is a significant biological cost. Flight is the most costly type of locomotion exemplified in the natural world. The energy expenditure required for flight increases proportionally with body size, which is often why flightlessness coincides with body mass. By reducing large pectoral muscles