Origin of the domestic dog to the domestic dog and modern wolf lineages. The gray wolf is thought to be ancestral to the domestic dog, however its relationship to "C. variabilis", and the genetic contribution of "C. variabilis" to the dog, is the subject of debate. The Zhokhov Island (8,700 YBP) and Aachim (1,700 YBP) canid haplotypes fell within the domestic dog clade, cluster with S805, and also share their haplotypes with – or are one mutation away from – the Tibetan wolf ("C. l. chanco") and the recently-extinct Japanese wolf ("C. l. hodophilax"). This may indicate that these canids retained the genetic signature of
Origin of the domestic dog matched to "Canis lupus familiaris" - dog, and confirms that the Oberkassel dog is a direct ancestor of today's dogs. The bodies were dated to 14,223 YBP. This implies that in Western Europe there were morphologically and genetically "modern" dogs in existence around 14,500 years ago. Later studies assigned more of the other animal bones to the dog until most of a skeleton could be assembled. The humans were a man aged 40 years and a woman aged 25 years. All three skeletal remains were found covered with large 20 cm thick basalt blocks and were sprayed with red hematite