Origin of the domestic dog to the domestic dog and modern wolf lineages. The gray wolf is thought to be ancestral to the domestic dog, however its relationship to "C. variabilis", and the genetic contribution of "C. variabilis" to the dog, is the subject of debate. The Zhokhov Island (8,700 YBP) and Aachim (1,700 YBP) canid haplotypes fell within the domestic dog clade, cluster with S805, and also share their haplotypes with – or are one mutation away from – the Tibetan wolf ("C. l. chanco") and the recently-extinct Japanese wolf ("C. l. hodophilax"). This may indicate that these canids retained the genetic signature of
Origin of the domestic dog had been calculated in an earlier study. Using the 7,000 YBP specimen and this mutation rate, the dog-wolf divergence time is estimated to have occurred 36,900-41,500 YBP and this is consistent with the timing found with the Taimyr-1 specimen in an earlier study. In 2018, a study looked at the y-chromosome male lineage of canines, including the ancient fossils of the Herxheim, Kirschbaum, and Newgrange dogs. The study identified six major dog yDNA haplogroups, of which two of these include the majority of modern dogs. The Newgrange dog fell into the most commonly occurring of these haplogroups. The two ancient