Digestive system of gastropods species that have undergone de-torsion, however, the oesophagus may open into the anterior of the stomach, which is therefore reversed from the usual gastropod arrangement. In "Tarebia granifera" the brood pouch is above the oesophagus. There is available an extensive rostrum on the anterior part of the oesophagus in all carnivorous gastropods. Some basal gastropod clades have oesophageal gland. In most species, the stomach itself is a relatively simple sac, and is the main site of digestion. In many herbivores, however, the hind part of the oesophagus is enlarged to form a crop, which, in terrestrial pulmonates, may even replace
Development of the digestive system anterior region (future urogenital sinus). The urorectal septum divides the two regions and breakdown of the cloacal membrane covering this area provides communication to the exterior for the anus and urogenital sinus. The upper part of the anal canal is derived from endoderm of the hindgut. The lower part (one-third) is derived from ectoderm around the proctodeum. Ectoderm, in the region of the proctodeum on the surface of part of the cloaca, proliferates and invaginates to create the anal pit. Subsequently, degeneration of the cloacal membrane establishes continuity between the upper and lower parts of the anal canal. Abnormalities in