Southern Ocean Southern Hemisphere, with the greatest concentration located on and around Antarctica. Four of the 18 penguin species live and breed on the mainland and its close offshore islands. Another four species live on the subantarctic islands. Emperor penguins have four overlapping layers of feathers, keeping them warm. They are the only Antarctic animal to breed during the winter. There are relatively few fish species in few families in the Southern Ocean. The most species-rich family are the snailfish (Liparidae), followed by the cod icefish (Nototheniidae) and eelpout (Zoarcidae). Together the snailfish, eelpouts and notothenioids (which includes cod icefish and several
Flightless bird free to increase in abundance until the population was limited by food and territory. A study looking at energy conservation and the evolution of flightlessness hypothesized intraspecific competition selected for a reduced individual energy expenditure, which is achieved by the loss of flight. Some flightless varieties of island birds are closely related to flying varieties, implying flight is a significant biological cost. Flight is the most costly type of locomotion exemplified in the natural world. The energy expenditure required for flight increases proportionally with body size, which is often why flightlessness coincides with body mass. By reducing large pectoral muscles