Origin of the domestic dog to the domestic dog and modern wolf lineages. The gray wolf is thought to be ancestral to the domestic dog, however its relationship to "C. variabilis", and the genetic contribution of "C. variabilis" to the dog, is the subject of debate. The Zhokhov Island (8,700 YBP) and Aachim (1,700 YBP) canid haplotypes fell within the domestic dog clade, cluster with S805, and also share their haplotypes with – or are one mutation away from – the Tibetan wolf ("C. l. chanco") and the recently-extinct Japanese wolf ("C. l. hodophilax"). This may indicate that these canids retained the genetic signature of
Dog training punishment are at risk of developing disturbances associated with the learned helplessness disorder. Punishment which is poorly coordinated with identifiable avoidance cues or response options, such as when punishment takes place long after the event, meet the criteria of inescapable trauma. Observational learning is the learning that occurs through observing the behavior of others. This form of learning does not need reinforcement to occur; instead, a model animal is required. While the model may not be intentionally trying to instill any particular behavior, many behaviors that are observed are remembered and imitated. The domestic dog is a social species and