Photosynthesis algae, photosynthesis takes place in organelles called chloroplasts. A typical plant cell contains about 10 to 100 chloroplasts. The chloroplast is enclosed by a membrane. This membrane is composed of a phospholipid inner membrane, a phospholipid outer membrane, and an intermembrane space. Enclosed by the membrane is an aqueous fluid called the stroma. Embedded within the stroma are stacks of thylakoids (grana), which are the site of photosynthesis. The thylakoids appear as flattened disks. The thylakoid itself is enclosed by the thylakoid membrane, and within the enclosed volume is a lumen or thylakoid space. Embedded in the thylakoid membrane are
Fractionation of carbon isotopes in oxygenic photosynthesis processes that selectively use one carbon isotope over the other, or "fractionate" carbon through kinetic or thermodynamic effects. Oxygenic photosynthesis takes place in plants and microorganisms through different chemical pathways, so various forms of organic material reflect different ratios of C isotopes. Understanding these variations in carbon fractionation across species is applied in isotope geochemistry and ecological isotope studies to understand biochemical processes, establish food chains, or model the carbon cycle through geological time. Carbon isotope fractionations are expressed in using delta notation of "δ"C ("delta thirteen C"), which is reported in parts per thousand (per mille, ‰). "δ"C is