Albugo occidentalis Albugo occidentalis Albugo occidentalis, the causal agent of Spinach White Rust, is an oomycete plant pathogen, although some discussions still treat it as a fungal organism. "Albugo occidentalis" is one of the most important spinach diseases in North America, found throughout the United States east of the rocky mountains. The economically important host of "Albugo occidentalis" is spinach (Spinacia oleracea); although the oomycete has also been reported to affect plants of the genus Chenopodium, the genus including the crop plant quinoa. This pathogen causes white rust or white blister of spinach. It is unrelated to the basidiomycete rusts biologically, but
Four causes after the ethologist Nikolaas Tinbergen and based on Aristotle's four causes, are complementary categories of explanations for animal behaviour. They are also commonly referred to as levels of analysis. The four questions are on function, what an adaptation does that is selected for in evolution; phylogeny, the evolutionary history of an organism, revealing its relationships to other species; mechanism, namely the proximate cause of a behaviour, such as the role of testosterone in aggression; and ontogeny, the development of an organism from egg to embryo to adult. Four causes The "four causes" are elements of an influential principle in Aristotelian