Rust (fungus) of the rust fungi "Cronartium ribicola", preventing the formation of basidiospores which infect the primary host. Although spores from white pines cannot infect other white pines, survival spores may overwinter on infected pines and reinfect "Ribes" spp. the following season. Infected tissue is removed from white pines and strict quarantines of "Ribes" spp. are maintained in high risk areas. "Puccinia graminis" is a macrocyclic heteroecious fungus that causes wheat stem rust disease. The repeating stage in this fungus occurs on wheat and not the alternate host, barberry. The repeating stage allows the disease to persist in wheat even though the
Stem rust studied the stem rust. Thirty years later it received its name, "Puccinia graminis", by Persoon, and in 1854 brothers Louis René and Charles Tulasne discovered the characteristic five-spore stage that is known to some stem rust species. The brothers were also able to make a connection between the red (urediniospore) and black (teliospore) spores as different stages within the same organism, but the rest of the stages remained unknown. Anton de Bary later conducted experiments to observe the beliefs of the European farmers regarding the relationship between the rust and barberry plants, and after successful attempts to connect the basidiospores