Marine life appeared in the form of fish, which live exclusively in water. Some of these evolved into amphibians which spend portions of their lives in water and portions on land. Other fish evolved into land mammals and subsequently returned to the ocean as seals, dolphins or whales. Plant forms such as kelp and algae grow in the water and are the basis for some underwater ecosystems. Plankton, and particularly phytoplankton, are key primary producers forming the general foundation of the ocean food chain. Marine invertebrates exhibit a wide range of modifications to survive in poorly oxygenated waters including breathing tubes (see
Marine biology on the shore. Many land animals also make much use of the shore and intertidal habitats. A subgroup of organisms in this habitat bores and grinds exposed rock through the process of bioerosion. Estuaries are also near shore and influenced by the tides. An estuary is a partially enclosed coastal body of water with one or more rivers or streams flowing into it and with a free connection to the open sea. Estuaries form a transition zone between freshwater river environments and saltwater maritime environments. They are subject both to marine influences—such as tides, waves, and the influx of saline