Albertosaurus older "Gorgosaurus libratus" (sometimes called "Albertosaurus libratus"; see below). These two species are the only described albertosaurines; other undescribed species may exist. Thomas Holtz found "Appalachiosaurus" to be an albertosaurine in 2004, but his more recent unpublished work locates it just outside Tyrannosauridae, in agreement with other authors. The other major subfamily of tyrannosaurids is the Tyrannosaurinae, which includes "Daspletosaurus", "Tarbosaurus" and "Tyrannosaurus". Compared with these robust tyrannosaurines, albertosaurines had slender builds, with proportionately smaller skulls and longer bones of the lower leg (tibia) and feet (metatarsals and phalanges). Below is the cladogram of the Tyrannosauridae based on the phylogenetic
Diandongosuchus The astragalus and calcaneum bones of the ankle fit together like a ball-and-socket, a feature that confirms "Diandongosuchus" as a pseudosuchian. Some of the phalanges or toe bones are missing in ZMNH M8770, but the metatarsals are present and have unique proportions among Triassic archosaurs in which the fourth is longer than the third. A phylogenetic analysis conducted by Li "et al." (2012) in the original description of "Diandongosuchus" showed that it was the most basal member of a clade called Poposauroidea, which includes mostly terrestrial pseudosuchians such as the bipedal "Poposaurus" and the sail-backed "Arizonasaurus". It was found to