Work of breathing When there is increased gas flow resistance, the optimal respiratory rate decreases. This work (generally during the inhalation phase) is stored as potential energy which is recovered during exhalation. A pressure difference is required to overcome the frictional resistance to gas flow due to viscosity, and to provide non-elastic components of movement of the airway tissues to accommodate pulmonary volume change. Total work done against non elastic forces is 35% of total. Work is defined as a force applied over a distance. The SI unit of work is the Joule, equivalent to a force of 1 Newton exerted along a
Rad (unit) in the latest NIST definition. Even where the standard medium is fully defined, the ionizing energy is often not precisely known. In 1940, British physicist Louis Harold Gray, who had been studying the effect of neutron damage on human tissue, together with William Valentine Mayneord and John Read published a paper in which a unit of measure, dubbed the ""gram roentgen"" (symbol: gr) defined as "that amount of neutron radiation which produces an increment in energy in unit volume of tissue equal to the increment of energy produced in unit volume of water by one roentgen of radiation" was proposed.