Digestive system of gastropods species that have undergone de-torsion, however, the oesophagus may open into the anterior of the stomach, which is therefore reversed from the usual gastropod arrangement. In "Tarebia granifera" the brood pouch is above the oesophagus. There is available an extensive rostrum on the anterior part of the oesophagus in all carnivorous gastropods. Some basal gastropod clades have oesophageal gland. In most species, the stomach itself is a relatively simple sac, and is the main site of digestion. In many herbivores, however, the hind part of the oesophagus is enlarged to form a crop, which, in terrestrial pulmonates, may even replace
Human digestive system the duodenum. (The enzymes that digest polysaccharides, by contrast, are primarily produced by the walls of the intestines.) The cells are filled with secretory granules containing the precursor digestive enzymes. The major proteases, the pancreatic enzymes which work on proteins, are trypsinogen and chymotrypsinogen. Elastase is also produced. Smaller amounts of lipase and amylase are secreted. The pancreas also secretes phospholipase A2, lysophospholipase, and cholesterol esterase. The precursor zymogens, are inactive variants of the enzymes; which avoids the onset of pancreatitis caused by autodegradation. Once released in the intestine, the enzyme enteropeptidase present in the intestinal mucosa activates trypsinogen by