Southern Ocean Southern Hemisphere, with the greatest concentration located on and around Antarctica. Four of the 18 penguin species live and breed on the mainland and its close offshore islands. Another four species live on the subantarctic islands. Emperor penguins have four overlapping layers of feathers, keeping them warm. They are the only Antarctic animal to breed during the winter. There are relatively few fish species in few families in the Southern Ocean. The most species-rich family are the snailfish (Liparidae), followed by the cod icefish (Nototheniidae) and eelpout (Zoarcidae). Together the snailfish, eelpouts and notothenioids (which includes cod icefish and several
Megafauna This is also the period when megafaunal flightless herbivorous gastornithid birds evolved in the Northern Hemisphere, while flightless paleognaths evolved to large size on Gondwanan land masses and Europe. Gastornithids and at least one lineage of flightless paleognath birds originated in Europe, both lineages dominating niches for large herbivores while mammals remained below 45 kg (in contrast with other landmasses like North America and Asia, which saw the earlier evolution of larger mammals) and were the largest European tetrapods in the Paleocene. Flightless paleognaths, termed ratites, have traditionally been viewed as representing a lineage separate from that of their small