Origin of the domestic dog to the domestic dog and modern wolf lineages. The gray wolf is thought to be ancestral to the domestic dog, however its relationship to "C. variabilis", and the genetic contribution of "C. variabilis" to the dog, is the subject of debate. The Zhokhov Island (8,700 YBP) and Aachim (1,700 YBP) canid haplotypes fell within the domestic dog clade, cluster with S805, and also share their haplotypes with – or are one mutation away from – the Tibetan wolf ("C. l. chanco") and the recently-extinct Japanese wolf ("C. l. hodophilax"). This may indicate that these canids retained the genetic signature of
Origin of the domestic dog result of a female wolf/male dog pairing. Haplogroup A: Includes 64-65% of dogs. Haplotypes of subclades a2–a6 are derived from post-domestication wolf–dog hybridization. Haplogroup B: Includes 22-23% of dogs. haplotypes of subclade b2 are derived from post-domestication wolf–dog hybridization. Haplogroup C: Includes 10-12% of dogs. Haplogroup D: Derived from post-domestication wolf–dog hybridization in subclade d1 (Scandinavia) and d2 (South-West Asia). The northern Scandinavian subclade d1 hybrid haplotypes originated 480-3,000 YBP and are found in all Sami-related breeds: Finnish Lapphund, Swedish Lapphund, Lapponian Herder, Jamthund, Norwegian Elkhound and Hällefors Elkhound. The maternal wolf sequence that contributed to them has not been