Origin of the domestic dog to the domestic dog and modern wolf lineages. The gray wolf is thought to be ancestral to the domestic dog, however its relationship to "C. variabilis", and the genetic contribution of "C. variabilis" to the dog, is the subject of debate. The Zhokhov Island (8,700 YBP) and Aachim (1,700 YBP) canid haplotypes fell within the domestic dog clade, cluster with S805, and also share their haplotypes with – or are one mutation away from – the Tibetan wolf ("C. l. chanco") and the recently-extinct Japanese wolf ("C. l. hodophilax"). This may indicate that these canids retained the genetic signature of
Origin of the domestic dog 2018, a study compared sequences of fossil North American dogs with fossil Siberian dogs and modern dogs. The study indicates that dogs entered North America from Siberia 4,500 years after humans did, were isolated for 9,000 years, and after contact with Europeans these no longer exist because they were replaced with Eurasian dogs. The pre-contact dogs exhibit a unique genetic signature that is also now gone, with their nearest genetic relatives being the arctic breed dogs. In 2017, a study showed that 9,000 YBP the domestic dog was present at what is now Zhokhov Island, arctic north-eastern Siberia, which was