Marine life appeared in the form of fish, which live exclusively in water. Some of these evolved into amphibians which spend portions of their lives in water and portions on land. Other fish evolved into land mammals and subsequently returned to the ocean as seals, dolphins or whales. Plant forms such as kelp and algae grow in the water and are the basis for some underwater ecosystems. Plankton, and particularly phytoplankton, are key primary producers forming the general foundation of the ocean food chain. Marine invertebrates exhibit a wide range of modifications to survive in poorly oxygenated waters including breathing tubes (see
Ecology of the San Francisco Estuary but by physiological tolerance to salinity and temperature. The Low Salinity Zone (LSZ) of the San Francisco Estuary constitutes a habitat for a suite of organisms that are specialized to survive in this unique confluence of terrestrial, freshwater, and marine influences. While there are many habitats with distinct ecologies that are part of the estuary (including marine, freshwater, intertidal marsh and benthic mudflat systems) each is linked to the LSZ by export and import of freshwater, nutrients, carbon, and organisms. The distribution and abundance of organisms in the LSZ is dependent upon both abiotic and biotic factors. Abiotic factors include