Origin of the domestic dog haplotype designated as S805 (28,000 YBP) from the Yana River was one mutation away from another haplotype S902 (8,000 YBP) that represents Clade A of the modern wolf and domestic dog lineages. Closely related to this haplotype was one that was found in the recently-extinct Japanese wolf. Several ancient haplotypes were oriented around S805, including "Canis c.f. variabilis" (360,000 YBP), Belgium (36,000 YBP – the "Goyet dog"), Belgium (30,000 YBP), and Konsteki, Russia (22,000 YBP). Given the position of the S805 haplotype on the phylogenetic tree, it may potentially represent a direct link from the progenitor (including "Canis c.f. variabilis")
Origin of the domestic dog a direct ancestor). The data from this study indicated a European origin for dogs that was estimated at 18,800–32,100 years ago based on the genetic relationship of 78% of the sampled dogs with ancient canid specimens found in Europe. The data supports the hypothesis that dog domestication preceded the emergence of agriculture and was initiated close to the Last Glacial Maximum when hunter-gatherers preyed on megafauna. The study found that three ancient Belgium canids (the 36,000 YBP "Goyet dog" cataloged as "Canis" species, along with Belgium 30,000 YBP and 26,000 years YBP cataloged as "Canis lupus") formed an ancient clade