Meadowsweet rust gall Meadowsweet rust gall Triphragmium ulmariae is a species of fungus in the Sphaerophragmiaceae family. It causes meadowsweet rust gall, which develops as a chemically induced swelling, arising from the lower surface of the meadowsweet leaves. This common gall is caused by the foliar 'rust' fungus "Triphragmium ulmariae" (DC.) Link, 1825 . The fungus attacks the petioles and / or midribs of the perennial meadowsweet ("Filipendula ulmaria") plant, a member of the family Rosaceae, causing swelling and distortion. Sori develop with bright orange spores. The rust's spores reach the meadowsweet via air movements and are spread from the galls that develop
Four causes after the ethologist Nikolaas Tinbergen and based on Aristotle's four causes, are complementary categories of explanations for animal behaviour. They are also commonly referred to as levels of analysis. The four questions are on function, what an adaptation does that is selected for in evolution; phylogeny, the evolutionary history of an organism, revealing its relationships to other species; mechanism, namely the proximate cause of a behaviour, such as the role of testosterone in aggression; and ontogeny, the development of an organism from egg to embryo to adult. Four causes The "four causes" are elements of an influential principle in Aristotelian