Holoaden Holoaden Holoaden (common name: highland frogs) is a small genus of craugastorid frogs endemic to southeastern Brazil. "Holoaden" is the type genus of subfamily Holoadeninae that was erected in 2008 and placed in family Strabomantidae, erected at the same time. Before that, it was in Brachycephalidae. However, later research has led the whole Strabomantidae, Holoadeninae included, to become included in Craugastoridae. This arrangement is not followed by all, and the AmphibiaWeb still keeps Holoadeninae, and consequently "Holoaden", in Strabomantidae. "Holoaden" grow to snout–vent length at most. Head is narrower than the body. Differentiated tympanic membrane and tympanic annulus are absent.
Inferior tympanic artery Inferior tympanic artery The inferior tympanic artery is a small branch of the ascending pharyngeal artery. It is a small branch which passes through a minute foramen in the petrous portion of the temporal bone which is called tympanic canaliculus or inferior tympanic canaliculus, in company with the tympanic branch of the glossopharyngeal nerve, to supply the medial wall of the tympanic cavity and anastomose with the other tympanic arteries. In the case of a missing or underdeveloped cervical ICA, the Inferior tympanic artery can provide collateral ICA circulation by reversing flow of the caroticotympanic artery (embryologic hyoid artery). This