Spermatophyte plants that have unenclosed, "naked seeds": The fifth extant division is the flowering plants, also known as angiosperms or magnoliophytes, the largest and most diverse group of spermatophytes. Angiosperms possess seeds enclosed in a fruit, unlike gymnosperms. In addition to the taxa listed above, the fossil record contains evidence of many extinct taxa of seed plants. The so-called "seed ferns" (Pteridospermae) were one of the earliest successful groups of land plants, and forests dominated by seed ferns were prevalent in the late Paleozoic. "Glossopteris" was the most prominent tree genus in the ancient southern supercontinent of Gondwana during the Permian
Evolutionary history of plants they were too scarce to impact the atmospheric composition until around . These organisms, although phylogenetically diverse, were probably small and simple, forming little more than an algal scum. However, evidence of the earliest land plants occurs much later at about 470Ma, in lower middle Ordovician rocks from Saudi Arabia and Gondwana in the form of spores with decay-resistant walls. These spores, known as cryptospores, were produced either singly (monads), in pairs (dyads) or groups of four (tetrads), and their microstructure resembles that of modern liverwort spores, suggesting they share an equivalent grade of organisation. Their walls contain sporopollenin –