Plant cell parenchyma cells, that covers the external surfaces of leaves, stems and roots. Several cell types may be present in the epidermis. Notable among these are the stomatal guard cells that control the rate of gas exchange between the plant and the atmosphere, glandular and clothing hairs or trichomes, and the root hairs of primary roots. In the shoot epidermis of most plants, only the guard cells have chloroplasts. Chloroplasts contain the green pigment chlorophyll which is needed for photosynthesis. The epidermal cells of aerial organs arise from the superficial layer of cells known as the "tunica" (L1 and L2 layers)
Evolution of photosynthesis Evolution of photosynthesis The evolution of photosynthesis refers to the origin and subsequent evolution of photosynthesis, the process by which light energy synthesizes sugars from carbon dioxide, releasing oxygen as a waste product. The process of photosynthesis was discovered by Jan Ingenhousz, a Dutch-born British physician and scientist, first publishing about it in 1779. The first photosynthetic organisms probably evolved early in the evolutionary history of life and most likely used reducing agents such as hydrogen or electrons, rather than water. There are three major metabolic pathways by which photosynthesis is carried out: C photosynthesis, C photosynthesis, and CAM photosynthesis.