Photosynthesis Earth's atmosphere, and supplies all of the organic compounds and most of the energy necessary for life on Earth. Although photosynthesis is performed differently by different species, the process always begins when energy from light is absorbed by proteins called reaction centres that contain green chlorophyll pigments. In plants, these proteins are held inside organelles called chloroplasts, which are most abundant in leaf cells, while in bacteria they are embedded in the plasma membrane. In these light-dependent reactions, some energy is used to strip electrons from suitable substances, such as water, producing oxygen gas. The hydrogen freed by the splitting
Fractionation of carbon isotopes in oxygenic photosynthesis processes that selectively use one carbon isotope over the other, or "fractionate" carbon through kinetic or thermodynamic effects. Oxygenic photosynthesis takes place in plants and microorganisms through different chemical pathways, so various forms of organic material reflect different ratios of C isotopes. Understanding these variations in carbon fractionation across species is applied in isotope geochemistry and ecological isotope studies to understand biochemical processes, establish food chains, or model the carbon cycle through geological time. Carbon isotope fractionations are expressed in using delta notation of "δ"C ("delta thirteen C"), which is reported in parts per thousand (per mille, ‰). "δ"C is