Marine life appeared in the form of fish, which live exclusively in water. Some of these evolved into amphibians which spend portions of their lives in water and portions on land. Other fish evolved into land mammals and subsequently returned to the ocean as seals, dolphins or whales. Plant forms such as kelp and algae grow in the water and are the basis for some underwater ecosystems. Plankton, and particularly phytoplankton, are key primary producers forming the general foundation of the ocean food chain. Marine invertebrates exhibit a wide range of modifications to survive in poorly oxygenated waters including breathing tubes (see
Aquatic ecosystem lagoons and mangrove swamps. In the deep water, hydrothermal vents may occur where chemosynthetic sulfur bacteria form the base of the food web. Classes of organisms found in marine ecosystems include brown algae, dinoflagellates, corals, cephalopods, echinoderms, and sharks. Fishes caught in marine ecosystems are the biggest source of commercial foods obtained from wild populations. Environmental problems concerning marine ecosystems include unsustainable exploitation of marine resources (for example overfishing of certain species), marine pollution, climate change, and building on coastal areas. Freshwater ecosystems cover 0.78% of the Earth's surface and inhabit 0.009% of its total water. They generate nearly 3%