Photosynthesis algae, photosynthesis takes place in organelles called chloroplasts. A typical plant cell contains about 10 to 100 chloroplasts. The chloroplast is enclosed by a membrane. This membrane is composed of a phospholipid inner membrane, a phospholipid outer membrane, and an intermembrane space. Enclosed by the membrane is an aqueous fluid called the stroma. Embedded within the stroma are stacks of thylakoids (grana), which are the site of photosynthesis. The thylakoids appear as flattened disks. The thylakoid itself is enclosed by the thylakoid membrane, and within the enclosed volume is a lumen or thylakoid space. Embedded in the thylakoid membrane are
Fractionation of carbon isotopes in oxygenic photosynthesis elevated O concentrations would increase the efficiency of photorespiration. Atmospheric CO on Earth decreased abruptly at a point between 32-25 million years ago. This gave a selective advantage to the evolution of the C4 pathway, which can limit photorespiration rate despite the reduced ambient CO. Today, C4 plants represent roughly 5% of plant biomass on Earth, but about 23% of terrestrial carbon fixation. Types of plants which use C4 photosynthesis include grasses and economically important crops, such as maize, sugar cane, millet, and sorghum. Isotopic fractionation is differs between C4 carbon fixation and C3, due to the spatial separation in