Henry Jones Shrapnell tympanic membrane is known as "Shrapnell's membrane". During the same year, Shrapnell published two other articles in the same journal, these being in regards to the function of the tympanic membrane and the nerves of the ear. In 1833, he published an article (again in the same journal) on the anatomy of the incus. Henry Jones Shrapnell Henry Jones Shrapnell (1792–1834) was an English anatomist. For a period of time during his career he was a colleague to Edward Jenner (1749-1823), creator of the vaccine for smallpox. Shrapnell is remembered for his pioneer work in otology. He was the first
Translocase of the inner membrane modified complex containing Tim12, on the outer surface of the TIM22 complex. Tim12 is membrane bound and thus may act as a linker molecule docking Tim9 and Tim10 to the face of the TIM22 complex. The carrier preprotein is then inserted into the inner mitochondrial membrane in a potential-dependent fashion. The membrane potential is necessary for both insertion of the precursor into the carrier translocase and lateral release of the protein into the lipid phase of the inner mitochondrial membrane, which completes protein translocation. However this membrane potential-dependent process takes place in absence of ATP-driven machinery. TIM17A; TIMM17A; TIMM17B; TIMM22;