Flightless bird that require a significant amount of overall metabolic energy, ratites decrease their basal metabolic rate and conserve energy. A study looking at the basal rates of birds found a significant correlation between low basal rate and pectoral muscle mass in kiwis. On the contrary, flightless penguins exude an intermediate basal rate. This is likely because penguins have well-developed pectoral muscles for hunting and diving in the water. For ground feeding birds, a cursorial lifestyle is more economical and allows for easier access to dietary requirements. Flying birds have different wing and feather structures that make flying easier, while flightless birds'
Evolution of birds more rigid ribcage with a carina and shoulders able to allow for a powerful upstroke, essential to sustained powered flight. Another improvement was the appearance of an alula, used to achieve better control of landing or flight at low speeds. They also had a more derived pygostyle, with a ploughshare-shaped end. An early example is "Yanornis". Many were coastal birds, strikingly resembling modern shorebirds, like "Ichthyornis", or ducks, like" Gansus". Some evolved as swimming hunters, like the Hesperornithiformes – a group of flightless divers resembling grebes and loons. While modern in most respects, most of these birds retained typical reptilian-like