Albertosaurus older "Gorgosaurus libratus" (sometimes called "Albertosaurus libratus"; see below). These two species are the only described albertosaurines; other undescribed species may exist. Thomas Holtz found "Appalachiosaurus" to be an albertosaurine in 2004, but his more recent unpublished work locates it just outside Tyrannosauridae, in agreement with other authors. The other major subfamily of tyrannosaurids is the Tyrannosaurinae, which includes "Daspletosaurus", "Tarbosaurus" and "Tyrannosaurus". Compared with these robust tyrannosaurines, albertosaurines had slender builds, with proportionately smaller skulls and longer bones of the lower leg (tibia) and feet (metatarsals and phalanges). Below is the cladogram of the Tyrannosauridae based on the phylogenetic
Ostromia addition, the ungual (claw) of the first digit of the hand is smaller than the corresponding first metacarpal, while in "Archaeopteryx" the claw is larger. The Haarlem specimen's metatarsals are also estimated to be proportionally longer than those of "Archaeopteryx" specimens. In addition, the Haarlem specimen shares several features with "Anchiornis". Most notably, they both have longitudinal furrows on the top and bottom sides of their manual phalanges (finger bones). While such structures can be a result of collapsed or broken bones (as is the case in several "Archaeopteryx" specimens), the straight, smooth edges of the furrows in "Ostromia" and