Plant cell parenchyma cells, that covers the external surfaces of leaves, stems and roots. Several cell types may be present in the epidermis. Notable among these are the stomatal guard cells that control the rate of gas exchange between the plant and the atmosphere, glandular and clothing hairs or trichomes, and the root hairs of primary roots. In the shoot epidermis of most plants, only the guard cells have chloroplasts. Chloroplasts contain the green pigment chlorophyll which is needed for photosynthesis. The epidermal cells of aerial organs arise from the superficial layer of cells known as the "tunica" (L1 and L2 layers)
Photosynthesis light, oxidize cytochrome f, chlorophyll a (and other pigments) will absorb another light, but will reduce this same oxidized cytochrome, stating the two light reactions are in series. In 1893, Charles Reid Barnes proposed two terms, "photosyntax" and "photosynthesis", for the biological process of "synthesis of complex carbon compounds out of carbonic acid, in the presence of chlorophyll, under the influence of light". Over time, the term "photosynthesis" came into common usage as the term of choice. Later discovery of anoxygenic photosynthetic bacteria and photophosphorylation necessitated redefinition of the term. After WWII at late 1940 at the University of California,