Marine life appeared in the form of fish, which live exclusively in water. Some of these evolved into amphibians which spend portions of their lives in water and portions on land. Other fish evolved into land mammals and subsequently returned to the ocean as seals, dolphins or whales. Plant forms such as kelp and algae grow in the water and are the basis for some underwater ecosystems. Plankton, and particularly phytoplankton, are key primary producers forming the general foundation of the ocean food chain. Marine invertebrates exhibit a wide range of modifications to survive in poorly oxygenated waters including breathing tubes (see
White-edge freshwater whipray In addition, the ampullae of Lorenzini of H. signifer are intermediate in size and structure between the large, complex ampullae of its marine relatives and the small, simple ampullae of potamotrygonid stingrays, which may be a consequence of varying salinities in its environment and demands that would place on electroreception. The white-edge freshwater whipray feeds mainly on small benthic organisms, including crustaceans, molluscs, and insect larvae. Reproduction is aplacental viviparous with the developing embryos sustained by maternally produced histotroph ("uterine milk"), as is the case in other stingrays. Newborns measure across; males and females mature sexually at and across respectively.