Horn of Africa polity was succeeded by the Sultanate of Ifat around 1285. Ifat was governed from its capital at Zeila in northern Somalia and was the easternmost district of the former Shewa Sultanate. The Adal Sultanate was a medieval multi-ethnic Muslim state centered in the Horn region. At its height, it controlled large parts of Somalia, Ethiopia, Djibouti and Eritrea. Many of the historic cities in the region, such as Amud, Maduna, Abasa, Berbera, Zeila and Harar, flourished during the kingdom's golden age. This period that left behind numerous courtyard houses, mosques, shrines and walled enclosures. Under the leadership of rulers such
Horn of Africa species total and about 90 species which are found exclusively in the region. Among endemic reptile genera, there are "Haackgreerius", "Haemodracon", "Ditypophis", "Pachycalamus" and "Aeluroglena". Half of these genera are uniquely found on Socotra. Unlike reptiles, amphibians are poorly represented in the region. There are about 100 species of freshwater fish in the Horn of Africa, about 10 of which are endemic. Among the endemic, the cave-dwelling Somali blind barb and the Somali cavefish can be found. It is estimated that about 5,000 species of vascular plants are found in the Horn, about half of which are endemic. Endemism is