Horn of Africa polity was succeeded by the Sultanate of Ifat around 1285. Ifat was governed from its capital at Zeila in northern Somalia and was the easternmost district of the former Shewa Sultanate. The Adal Sultanate was a medieval multi-ethnic Muslim state centered in the Horn region. At its height, it controlled large parts of Somalia, Ethiopia, Djibouti and Eritrea. Many of the historic cities in the region, such as Amud, Maduna, Abasa, Berbera, Zeila and Harar, flourished during the kingdom's golden age. This period that left behind numerous courtyard houses, mosques, shrines and walled enclosures. Under the leadership of rulers such
Horn of Africa pygmy gerbil, the ammodile, and the Speke's pectinator. The Grevy's zebra is the unique wild equid of the region. There are predators such as spotted hyena, striped hyena and African leopard. The endangered painted hunting dog had populations in the Horn of Africa, but pressures from human exploitation of habitat along with warfare have reduced or extirpated this canid in this region. Some important bird species of the Horn are the black boubou, the golden-winged grosbeak, the Warsangli linnet, and the Djibouti francolin. The Horn of Africa holds more endemic reptiles than any other region in Africa, with over 285