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The proton motive force arises from a combination of the concentration gradient of protons and a voltage gradient across the membrane (the protons are positively charged, so an imbalance of protons means that the side of the membrane with more protons has a positive charge relative to the side with fewer protons).
How Typhoons Occur. Researchers are continuously working toward and discovering new information on the phenomena of tropical cyclones as there is still much regarding the system that is unknown. Typhoons occur when a rough weather wave, using the Earth's rotation, begins to rotate (also known as the Coriolis effect).