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Dominating this era was Titanoboa, the undisputed largest snake in the history of the world. Most of the fossil record of ancient snakes is comprised of vertebrae like the one that launched the Titanoboa investigation.
If an official measurement confirms this, it will be the longest snake on record. Currently, the Guinness World Record-holder is Medusa, a reticulated python who resides in Kansas City, Mo. When she was measured in 2011, she was 25 feet 2 inches long, roughly a little longer than a mid-sized car.