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Dominating this era was Titanoboa, the undisputed largest snake in the history of the world. Most of the fossil record of ancient snakes is comprised of vertebrae like the one that launched the Titanoboa investigation.
As a side note â the largest snake species to ever exist on the Earth (as far as we know currently), was the titanoboa, which lived somewhere around 58-60 million years ago. Titanoboa cerrejonensis is estimated to have regularly reached sizes of 40-50 feet in length, and 2000-2500 lbs.