Substrate is very often a big debate in keeping tortoises. I have Russian tortoises, I try to mimic their natural habitat as closely as possible.Since they are an arid species, I use a 50/50 mixture of playsand and topsoil deep enough for the tortoise to burrow.ubstrate is very often a big debate in keeping tortoises. I have Russian tortoises, I try to mimic their natural habitat as closely as possible.
There are four major types of headache: vascular, muscle contraction (tension), traction, and inflammatory. The most common type of vascular headache is a migraine. Migraine headaches are usually characterized by severe pain on one or both sides of the head, nasuea, and, at times, disturbed vision. Women are more likely than men to have migraine headaches.
pulmonary vascular resistance the vascular resistance of the pulmonary circulation; the difference between the mean pulmonary arterial pressure and the left atrial filling pressure divided by the cardiac output. Called also total pulmonary vascular resistance. total peripheral resistance the vascular resistance of the systemic circulation: the difference between the mean arterial pressure and central venous pressure divided by the cardiac output. total pulmonary resistance (total pulmonary vascular resistance) pulmonary vascular resistance.