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In photosynthesis, the light-dependent reactions take place on the thylakoid membranes. The inside of the thylakoid membrane is called the lumen, and outside the thylakoid membrane is the stroma, where the light-independent reactions take place.The thylakoid membrane contains some integral membrane protein complexes that catalyze the light reactions.There are four major protein complexes in the thylakoid membrane: Photosystem II (PSII), Cytochrome b6f complex, Photosystem I (PSI), and ATP synthase.he thylakoid membrane contains some integral membrane protein complexes that catalyze the light reactions. There are four major protein complexes in the thylakoid membrane: Photosystem II (PSII), Cytochrome b6f complex, Photosystem I (PSI), and ATP synthase.
The light-dependent reaction can only occur in the light because light supplies the energy needed for the reaction. In this stage of photosynthesis, light energy is converted into chemical energy (ATP) and 'reducing power' (NADPH) which are then used in the light-independent reaction.Also in the light-dependent reaction, water is split to provide the hydrogen needed to make NADPH.owever, it needs the products of the light-dependent reaction (ATP and NADPH), so it cannot continue indefinitely without it. In the light-independent reaction, the ATP and NADPH produced by the light-dependent reaction are used to convert carbon dioxide into carbohydrate, e.g., glucose.