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Our towels were never able to dry out due to the humidity and so have been hanging damp for this past week and all smelled. I just finished trying the method suggested in this post and it worked great. I washed my towels twice, once in vinegar (1 cup) and once in baking soda (1/2 cup).
Finally, some dry environments have a musty odor. One cause of this odor that I call the Antique Shop Odor is degradation of paper products and very old wood. Old waxes and polishes could also contribute. I am not aware of publications that discuss the specific compounds that cause these odors.