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Biochemistry of Dog Coat Color. The color of a dogâs coat is determined by relative levels of two pigments: eumelanin, which can be either brown or black, and pheomelanin, which is yellow to orange or red. Both pigments are generated by metabolism of tyrosine. A schematic diagram of the pathway is below. Schematic of the melanin synthesis pathways.
Coat color is determined by two genes that have two alleles each. One gene, called E, determines whether the dogâs fur is dark or light, with the dark allele (E) dominant to the light allele (e). The other gene, called B, determines whether the dark fur will be black or chocolate, with the black allele (B) dominant to the chocolate allele (b).