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Biochemistry of Dog Coat Color. The color of a dogâs coat is determined by relative levels of two pigments: eumelanin, which can be either brown or black, and pheomelanin, which is yellow to orange or red. Both pigments are generated by metabolism of tyrosine. A schematic diagram of the pathway is below. Schematic of the melanin synthesis pathways.
Liver is recessive (b), so a dog needs two copies of the allele in order to be liver. If a dog is bb on the B locus then any and all black in its coat will become a brownish colour. Bb and BB have no effect. There are a number of different versions of the b allele, but all of them result in the same coat colour.