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In tetrapods, the carpus is the sole cluster of bones in the wrist between the radius and ulna and the metacarpus. The bones of the carpus do not belong to individual fingers (or toes in quadrupeds), whereas those of the metacarpus do. The corresponding part of the foot is the tarsus. The carpal bones allow the wrist to move and rotate vertically.
The distal row of the tarsus has four bones, three cuneiforms and a cuboid. In the former the calcic scales and crystals are distinct and quite as in L. tigrinum; in the latter they are cuboid, irregular. The external border of the foot lay next to the plate, as is shown by the articulation of the cuboid.