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Since the initial observation that GTP reacted with a soluble protein factor(s), present in an ammonium chloride extract of E. coli ribosomes, to form a complex that was retained on a nitrocel- lulose filter (11), it has been shown that both transfer factors, Ts and Tu, are required to form the GTP complex (6,8).
Peptidyl transferase is also involved in the termination reaction of protein synthesis and hydrolyzes the ester bond between the peptide chain and tRNA at the P site of the ribosome (see Release Factor).