Fossils of isolated land plant spores (Heckman et al, 2001) suggest land plants may have been around as long as 475 million years ago; Fossils of land plant spores within plant fragments (Wellman et al., 2003) date back to 425 million years ago ( these were found in Oman and appear to be liverwort-like, although elaters are noticeably absent!)
The noun PUPILLARY REFLEX has 1 sense: 1. reflex contraction of the sphincter muscle of the iris in response to a bright light (or certain drugs) causing the pupil to become smaller. Familiarity information: PUPILLARY REFLEX used as a noun is very rare. Dictionary entry details.
There are 6 muscles around the eye that are responsible for eye movement. They are called the superior rectus, inferior rectus, lateral rectus, medial rectus, superior obliqu â¦ e and inferior oblique.There are two muscles that affect the iris (color of the eye), they are the iris sphincter and iris dilator muscles.   Extrinsic muscles are voluntary muscles,external to the eye, that control the direction the eye moves in e.g looking left or right.   Intrinsic muscles are wit â¦ hin the eye, such as those that control the size of the pupil.