Fossils of isolated land plant spores (Heckman et al, 2001) suggest land plants may have been around as long as 475 million years ago; Fossils of land plant spores within plant fragments (Wellman et al., 2003) date back to 425 million years ago ( these were found in Oman and appear to be liverwort-like, although elaters are noticeably absent!)
The noun PUPILLARY REFLEX has 1 sense: 1. reflex contraction of the sphincter muscle of the iris in response to a bright light (or certain drugs) causing the pupil to become smaller. Familiarity information: PUPILLARY REFLEX used as a noun is very rare. Dictionary entry details.
Pupillary reflex. In medicine, there are two pupillary reflexes 1  -. The pupillary light reflex is the reduction of pupil size in response to light. 2  The pupillary accommodation reflex is the reduction of pupil size in response to an object coming close to the eye.he optic nerve is responsible for the afferent limb of the pupillary reflex, or in other words, senses the incoming light. The oculomotor nerve is responsible for the efferent limb of the pupillary reflex; in other words, it drives the muscles that constrict the pupil.