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One possibility is that it is a learned behavior that evolved from kiss feeding, the process by which mothers in some cultures feed their babies by passing masticated food from mouth-to-mouth. However, there are some contemporary indigenous cultures which still practice kiss feeding, but not social or erotic kissing.
A kiss on the forehead is a sweet, innocent way of saying he cares about you and you matter to him. There is a gallant, protective meaning behind a forehead kiss-but that doesn't necessarily mean it is romantic. However, if it was at a party it is likely he has romantic intentions, and is working his courage up slowly.