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American Box turtles are omnivores and will enjoy fruits, fungi, veggies, greens, mollusks, worms and insects. From research with wild turtles, we know they eat plants for almost half the diet and animal foods for a little bit over half the diet.
Essentially, if a box turtle can catch it, it will eat it. The rest of the box turtle's diet is composed plant matter. A large part of which is fungi (mushrooms etc). Moss, berries and grass are also eaten. Box turtles also consume stones and sand.