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WPA is a wireless encryption format and the passphrase is in the wireless security section of your router. Here is a link to one of the Sticky topics at the top of this forkum that should be of assistance.
Unless you are referring to the drivers, but it's doubtful that would require a key. So most likely the key you are referring to is the one used in the router and the one that's needed for the client to connect. In any case, you shouldn't use WEP or WPA. Instead, if possible use WPA2-PSK [AES]. WEP is hackable and WPA isn't as strong as WPA2. The Key or Passphrase should be a randomly generated 'password' that allows wireless clients, like your Desktop that you want to connect wirelessly, access to the router.