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Homeobox gene: A gene containing a short DNA sequence of about 180 base pairs referred to as a homeobox. Homeobox genes encode proteins that bind and regulate the expression of DNA in multicellular organisms.
VAHOX1, a homeobox gene from tomato, shows phloem specific expression during secondary growth and is therefore a candidate gene playing a role in phloem specification from the vascular cambium. It is known that the sieve element (SE) and companion cell (CC) arise from an unequal division of a common âphloem mother cell.â This mother cell may be found in the procambium in the case of primary phloem or in the vascular cambium in secondary phloem. The SE then undergoes a âpartial programmed cell death.â This highly selective degradation of cellular organalles eliminates the vacuole, cytoskeleton, ribosomes, Golgi bodies and nucleus.