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Homeobox gene: A gene containing a short DNA sequence of about 180 base pairs referred to as a homeobox. Homeobox genes encode proteins that bind and regulate the expression of DNA in multicellular organisms.
The homeobox encodes a 60-amino acid protein segment called the homeodomain, which recognizes and binds to promoters in the DNA of its target genes. Complete control over transcription, and sometimes binding, is dependent on interactions between TFs, so activation often depends upon the presence of another TF.