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In photosynthesis, the light-dependent reactions take place on the thylakoid membranes. The inside of the thylakoid membrane is called the lumen, and outside the thylakoid membrane is the stroma, where the light-independent reactions take place. The thylakoid membrane contains some integral membrane protein complexes that catalyze the light reactions. There are four major protein complexes in the thylakoid membrane: Photosystem II (PSII), Cytochrome b6f complex, Photosystem I (PSI), and ATP synthase.
Atp synthesis in organelles. 1  1. ATP synthesis in organelles Michael Poole. 2  2. Chloroplast and Mitochondria. 3  3. Chloroplast are Adapted to allow photosynthesisâ¢ The process of photosynthesis consists of two sets of chemical reactions called the light reaction and the dark reaction. The light reaction takes place in the grana. The dark reaction takes place in the stroma.â¢ Chloroplasts contain both DNA and ribosomes.