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In photosynthesis, the light-dependent reactions take place on the thylakoid membranes. The inside of the thylakoid membrane is called the lumen, and outside the thylakoid membrane is the stroma, where the light-independent reactions take place. The thylakoid membrane contains some integral membrane protein complexes that catalyze the light reactions. There are four major protein complexes in the thylakoid membrane: Photosystem II (PSII), Cytochrome b6f complex, Photosystem I (PSI), and ATP synthase.
There are two theories as to how reactions occur: 1  In the collision theory, it is thought that reactions occur because molecules collide; the faster they collide, the faster the reaction occurs. 2  In the transition state theory, reactants are thought to form bonds and then break bonds until they form products.