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In photosynthesis, the light-dependent reactions take place on the thylakoid membranes. The inside of the thylakoid membrane is called the lumen, and outside the thylakoid membrane is the stroma, where the light-independent reactions take place. The thylakoid membrane contains some integral membrane protein complexes that catalyze the light reactions. There are four major protein complexes in the thylakoid membrane: Photosystem II (PSII), Cytochrome b6f complex, Photosystem I (PSI), and ATP synthase.
The light reactions, which convert the energy in light into chemical energy, take place within the thylakoid membranes of the chloroplasts, whereas the dark reactions, which use that chemical energy to fix CO 2 into organic molecules, take place in the stroma of the chloroplast.