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In photosynthesis, the light-dependent reactions take place on the thylakoid membranes. The inside of the thylakoid membrane is called the lumen, and outside the thylakoid membrane is the stroma, where the light-independent reactions take place. The thylakoid membrane contains some integral membrane protein complexes that catalyze the light reactions. There are four major protein complexes in the thylakoid membrane: Photosystem II (PSII), Cytochrome b6f complex, Photosystem I (PSI), and ATP synthase.
Rating Newest Oldest. Best Answer: Light reactions take place on the thykaloid membranes of the chloroplast. Light reaction requires light and water, and produces ATP and Oxygen. Dark reactions take place in the stroma (fluid that fills the chloroplast). It uses the ATP produced in light reactions to make glucose. Source(s):