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Cholecystokinin is secreted by I-cells in the small intestine and induces contraction of the gall bladder, relaxes the sphincter of Oddi, reduces of gastric acid secretion, increases bile acid production in the liver, delays gastric emptying, and induces digestive enzyme production in the pancreas.
Contraction of the muscularis pushes bile out of the gallbladder and into the cystic duct. Surrounding the muscularis is a thin layer of fibrous connective tissue that helps to reinforce and strengthen the wall of the gallbladder.