Medical Definition of RED NUCLEUS. : a nucleus of gray matter in the tegmentum of the midbrain on each side of the middle line that receives fibers from the cerebellum of the opposite side by way of the superior cerebellar peduncle and gives rise to fibers of the rubrospinal tract of the opposite side. a nucleus of gray matter in the tegmentum of the midbrain on each side of the middle line that receives fibers from the cerebellum of the opposite side by way of the superior cerebellar peduncle and gives rise to fibers of the rubrospinal tract of the opposite side.
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