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Medical Definition of histone. : any of various simple water-soluble proteins that are rich in the basic amino acids lysine and arginine and are complexed with DNA in the nucleosomes of eukaryotic chromatin.
Proteins constitute over 80% of the total mass of nuclear material, while DNA forms 15 to 20%. There are 5 histone proteins associated with DNA Two molecules each of histones H2A, H2B, H3 and H4 form a histone core. DNA is wound around the histone cores-two turns of DNA around each. core form a nucleosome.