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Medical Definition of histone. : any of various simple water-soluble proteins that are rich in the basic amino acids lysine and arginine and are complexed with DNA in the nucleosomes of eukaryotic chromatin.
A histone octamer is the eight protein complex found at the center of a nucleosome core particle.It consists of two copies of each of the four core histone proteins (H2A, H2B, H3 and H4).s a result of the rotation, two ends of histones involved in DNA binding of the crescent shape H3-H4 are equivalent, yet they organize different stretches of DNA. The H2A-H2B dimer also folds similarly. The H3 2 -H4 2 tetramer is wrapped with DNA around it as a first step of nucleosome formation.