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Lord Edric Dayne tells Arya Stark that Wylla is Jon's mother. He recounts to Arya Stark that he was a milk-brother to Jon Snow since Edric was also nursed as a baby by Wylla. He also claims they are not blood brothers either.
The theory goes that Jon is not Eddard Stark's son, but his nephew, and that his true mother is Ned's sister Lyanna. What's more, his father is Rhaegar Targaryen, the silver-haired prince who incited Robert's Rebellion by abducting Lyanna.