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A grain is a unit of measurement of mass that is nominally based upon the mass of a single seed of a cereal. A seed is a small embryonic plant enclosed in a covering called the seed coat, usually with some stored food.
The main differences between the wild forms of wheat and domesticated wheat are that domesticated forms have larger seeds with hulls and a non-shattering rachis. When wild wheat is ripe, the rachis--the stem that keeps the wheat shafts together--shatters so that the seeds can disperse themselves.