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The Entangled Bank. Definition: Spindle fibers are aggregates of microtubules that move chromosomes during cell division. During prophase of mitosis, spindle fibers form at opposite poles of the cell. During metaphase, spindle fibers extend from the cell poles toward the midpoint of the cell known as the metaphase plate.Chromosomes are held at the metaphase plate by the equal forces of the spindle fibers pushing on the centromeres of the chromosomes.In anaphase, spindle fibers pull sister chromatids toward the spindle poles. Spindle fibers not connected to chromatids lengthen and elongate the cell.uring metaphase, spindle fibers extend from the cell poles toward the midpoint of the cell known as the metaphase plate. Chromosomes are held at the metaphase plate by the equal forces of the spindle fibers pushing on the centromeres of the chromosomes.
Textiles and manufacturing. 1  Spindle (textiles), a device to spin fibers into thread. 2  Spindle (tool), a rotating axis of the machine, which often has a shaft at its heart. 3  Segmented spindle, a spindle consisting of discrete elements that can be assembled and disassembled on the fly.