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Definition of evolve for English Language Learners. : to change or develop slowly often into a better, more complex, or more advanced state : to develop by a process of evolution.
With all the physical forces at work in an estuary (the flow of rivers; the deposition of sediments; the ebb and flow of tides) conditions are constantly changing. Species that evolve in estuaries are therefore adapted to tremendous variability and extreme conditions in their environment.