As a group, titanosaurs were high browsers, eating leaves and branches from tall trees. This is in contrast to other herbivores like the Ceratopsia and Ankylosauria, which were low browsers, eating lower vegetation like shrubs. As a general rule, high browsers can eat lower-growing plants, but low browsers cannot eat from tall trees. Like the earlier sauropods (e.g. "Diplodocus") titanosaurs had peg-like teeth and no grinders. This means they used gastroliths to grind their food, and had a huge vat-like stomach where bacteria decomposed it. 