The band's lineup was the same for "The Bends". Thom Yorke and Ed O'Brien played rhythm and auxiliary guitar; Thom sang and Ed sang backup. Phil Selway played drums and percussion. Colin Greenwood played bass guitar. Radiohead's lead guitarist was Jonny Greenwood, Colin's younger brother. Jonny was also younger than any of the other members. Jonny was the only member of the band who did not graduate from university. He was also the only member who had attended music school. Jonny had had to leave university after a few months to join Radiohead on a tour when they became popular. But he played in an orchestra when he was young, and he had a knowledge of classical and experimental music and jazz. Thus, Jonny became the band's musical expert. At this stage he already played many instruments, such as keyboard.