Dated to 3.6 million years ag,o they were the oldest known evidence of hominin bipedalism. Now older evidence has been found, such as the "Ardipithecus ramidus" fossils. The footprints and skeletal structure excavated at Laetoli were clear evidence that bipedalism evolved before enlarged brains in hominids. Although it is debated, it is believed the three individuals who made these footprints belonged to the species "Australopithecus afarensis". Together with footprints were other hominin and animal skeletal remains and Acheulean artifacts (hand-axes).