Other experimental observations can be explained by combining the overall expansion of space with nuclear physics and atomic physics. As the Universe expands, the energy density of the electromagnetic radiation decreases more quickly than does that of matter, since the energy of a photon decreases with its wavelength. Thus, although the energy density of the Universe is now dominated by matter, it was once dominated by radiation; poetically speaking, all was light. As the Universe expanded, its energy density decreased and it became cooler; as it did so, the elementary particles of matter could associate stably into ever larger combinations. Thus, in the early part of the matter-dominated era, stable protons and neutrons formed, which then associated into atomic nuclei. At this stage, the matter in the Universe was mainly a hot, dense plasma of negative electrons, neutral neutrinos and positive nuclei. Nuclear reactions among the nuclei led to the present abundances of the lighter nuclei, particularly hydrogen, deuterium, and helium. Eventually, the electrons and nuclei combined to form stable atoms, which are transparent to most wavelengths of radiation; at this point, the radiation decoupled from the matter, forming the ubiquitous, isotropic background of microwave radiation observed today.