According to Macrobius, Aphroditus was represented in Cyprus with a beard and phallus, wearing female clothing and holding a scepter. At sacrifices to the god, worshipers cross-dressed, men wearing women's clothing and women dressed in men's clothing. The cult spread from Cyprus into southern Asia Minor, eventually reaching mainland Greece around the late 5th century BC. In Athens, the iconography of the god took on the "anasyromenos" pose, a female figure lifting her dress to reveal an erect phallus, a gesture that had apotropaic qualities, much like the god Priapus. By the Hellenistic period, the cult and votives had spread throughout Asia Minor, Greece, and Italy.