The Nobel Prize in Physiology or Medicine 2002 was awarded jointly to Sydney Brenner, H. Robert Horvitz and John E. Sulston "for their discoveries concerning genetic regulation of organ development and programmed cell death'." Sydney Brenner, Sir John Sulston and Robert Horvitz received the Nobel Prize for their work on the genome and cells of the roundworm "Caenorhabditis elegans". C. elegans was the first animal to have its complete genome sequenced.