Later, four fossil skeletons were found, showing a species primitive in its skull and upper body but with relatively advanced spines and lower limbs, providing greater mobility. They are now thought to represent a stage soon after the transition between "Australopithecus" and "Homo erectus", and have been dated at 1.8 million years before the present. The assemblage includes one of the largest Pleistocene Homo mandibles (D2600), one of the smallest Lower Pleistocene mandibles (D211), a nearly complete sub-adult (D2735), and a specimen without teeth (D3900).