Many nearly complete fossil coelurosaurians are known from the late Jurassic. "Archaeopteryx" is known from Solnhofen limestone at 155-150 million years ago (mya). "Ornitholestes", the troodontid WDC DML 110, "Coelurus fragilis" and "Tanycolagreus topwilsoni" are all known from the Morrison Formation in Wyoming at about 150 mya. "Epidendrosaurus" and "Pedopenna" are known from the Daohugou Beds in China, whose age is still being debated, but may be about 160 Ma or 145 mya.