Fossil pterodactyloids appear in the Upper Jurassic. They were short-tailed pterosaurs, suggesting that they had more sophisticated control of their flight, which no doubt gave them some advantages. 2–300 specimens of 17 different species of pterosaur has been found at Solnhofen from eight different genera. They include the earliest examples of "Pterodactylus", and "Germanodactylus", a genus which is also found in England and China. "Ctenochasma", also from Solnhofen, had a comb of 260 thin teeth showing it to be a filter-feeder that may have waded or swam in the water. There were several other genera with similar life-styles.