Erinaceids are generally shrew-like in form, with long snouts and short tails, but they are much larger than shrews. All but one species have five toes in each foot (in some cases with strong claws for digging) and they have large eyes and ears. Hedgehogs have hair modified into sharp spines to form a protective covering over the upper body and flanks, while gymnures have only normal hair. Most species have anal scent glands, but these are far better developed in gymnures, which can have a powerful odor.