In a tree farm, three musically inclined chipmunks, Alvin (Justin Long), the mischievous troublemaker, Simon (Matthew Gray Gubler), the smart one in the trio, and Theodore (Jesse McCartney) the chubby sweethearted chipmunk, find their tree cut down and are transported to Los Angeles. Once there, they meet frustrated songwriter David Seville (Jason Lee), and despite a poor house-wrecking first impression, they impress him with their singing talent. Seeing the opportunity for success, both human and chipmunks make a pact for them to sing his songs. While that ambition proves a frustrating struggle with the difficult trio, the dream does come true after all. However, that success presents its own trials as their unscrupulous record executive, Ian Hawke (David Cross), plans to break up this family to exploit the boys. Can Dave and the Chipmunks discover what they really value amid the superficial glamour around them?