(S (S (S (VP (TO To) (VP (VB compare) (PP (IN against) (BASENP (DT a) (JJ second) (JJ large) (NN species)))))) (COMMA ,) (S (NP (BASENP (NNP Burda)) (CC and) (BASENP (PRPD his) (NN team))) (VP (VP (VBD analyzed) (BASENP (NN data))) (PP (IN on) (NP (BASENP (NN 2,974) (NN deer)) (VP (VBN studied) (PP (IN through) (NP (NP (BASENP (NN photography)) (COMMA ,) (BASENP (JJ direct) (NN observation))) (CONJP (COMMA ,) (CC and)) (BASENP (NN snow) (NNS imprints)))))))))) (PERIOD .))