(S (S (NP (NP (BASENP (DT The) (NN drug)) (VP (VBN called) (S (BASENP (NN PA-457))))) (PRN (COMMA ,) (SBAR (WHNP (WDT which)) (S (VP (VP (VBZ is) (VP (VBN derived) (PP (IN from) (BASENP (JJ betulinic) (NN acid))))) (CC and) (VP (VBZ comes) (PP (IN from) (NP (BASENP (NNS by-products)) (PP (IN from) (BASENP (NN paper) (NNS manufacturers))))))))) (COMMA ,))) (VP (VBZ is) (VP (VBG being) (VP (VP (VBN manufactured) (CC and) (VBN developed)) (PP (IN by) (BASENP (NNP Maryland) (NN drug) (NN manufacturer) (NNP Panacos) (NNPS Pharmaceuticals))))))) (PERIOD .))