(S (S (BASENP (DT The) (NN study)) (VP (VBD found) (NP (BASENP (NN evidence)) (PP (IN of) (NP (BASENP (JJ polar) (NN bear) (NN hybridization)) (PP (IN with) (NP (BASENP (DT a) (JJ now-extinct) (NN population)) (PP (IN of) (NP (BASENP (JJ brown) (NNS bears)) (SBAR (WHNP (WDT that)) (S (VP (VP (VBD lived) (PP (IN in) (BASENP (NNP Ireland) (CC and) (NNP Britain)))) (PP (IN during) (BASENP (DT the) (JJ last) (NN ice) (NN age))))))))))))))) (PERIOD .))