In the 6 - vs. 1-portion post-intervention comparisons , carotenoids increased in serum , HDL2 and particularly HDL3 , ( - carotene , p = 0.008 ; - cryptoxanthin , p = 0.042 ; lutein , p = 0.012 ; lycopene , p = 0.016 ) , as did the activities of PON-1 and LCAT in HDL3 ( p = 0.006 and 0.044 , respectively ) .