The perfective represented by "nu," "tsu" and "ri" and the past tense represented by "ki," "shi" and "keri" were obsolete and the perfect aspect "tari" developed into the general past tense.
完了を表す「ぬ」「つ」「り」、過去の時制を表す「き」「し」「けり」は廃れていき、代わりに完了相「たり」が一般的な過去時制へと発展していった。