On the most basic level, Guji are shrines' main representatives, Gonguji are deputy representatives, Negi are assistants to the Guji, and Gonnegi are general clerical workers.
簡単にいえば、宮司は神社の代表者、権宮司は副代表、禰宜は宮司の補佐役、権禰宜は一般職員である。