Sanuki Province Kozai Clan: Tanba Province Shugodai; Noto Province Yusa Clan: Kawachi Province Shugodai; etc.) increasingly often, and as the positions became hereditary, they replaced the Shugo as effective rulers.
讃岐国香西氏→丹波国守護代、能登国遊佐氏→河内国守護代など）ことが多くなり、次第に世襲化していくと、守護に代わって実質的統治者になっていった。