Naotaka encouraged the domain population to move and to form clusters and renamed the domain Kujira but died on October 8, 1865, succeeded by Naochika NAGAI.
直壮は領民の移住や集住を奨励し、藩名も正式に櫛羅藩と改めたが、慶応元年（1865年）8月19日に死去し、後を永井直哉が継ぐ。