Additionally, being influenced by the tantora of Shakta Hinduism and Shakti belief, Anunttara-yoga was seen characteristically in the late-stage Mikkyo, which was a training of coalescence (sex) between male principles (mind, wisdom, upaya and Vajradhatu) and female principles (body, feeling, Hannya and Garbhadhatu), so that there appeared joyful Buddha (歓喜仏) which showed that the male Buddha (male principle) and female Buddha (female principle) had sexual intercourse.
また、ヒンドゥー教シャークタ派のタントラやシャクティ（性力）信仰から影響を受け始めて、男性原理（精神・智・方便・金剛界）と女性原理（肉体・感・般若・胎蔵界）との合体（性交）を修行する無上瑜伽（性的ヨガ）も後期密教の特徴であり、男尊（男性原理）と女尊（女性原理）が性交する歓喜仏も多数登場した。