The earliest inhabitants of what is now Rwanda were the Twa, a group of aboriginal pygmy hunter-gatherers who settled the area between 8000 and 3000 BC and remain in the country today. They were followed between 700 BC and AD 1500 by a number of Bantu groups, including the Hutu and Tutsi, who began clearing forests for agriculture. According to oral history, the Kingdom of Rwanda was founded in the 14th century on the shores of Lake Muhazi, around  east of modern Kigali. The early kingdom included Kigali but it was a small state at this point in its history with larger and more powerful neighbours, Bugesera and Gisaka. A member of the Gisaka dynasty killed Rwanda's king Ruganzu I Bwimba in the 16th century, but Ruganzu's son Cyilima I Rugwe fought back with help from Bugesera and was able to expand Rwanda's territory. In the late 16th or early 17th century, the kingdom of Rwanda was invaded from the north by the Banyoro of modern-day Uganda. The king was forced to flee westward, leaving Kigali and eastern Rwanda in the hands of Bugesera and Gisaka. The formation of a new Rwandan dynasty in the 17th century by the ''mwami'' (king), Ruganzu II Ndoli, followed by eastward invasions and the conquest of Bugesera, marked the beginning of the Rwandan kingdom's dominance in the area. The capital of the kingdom was at Nyanza, in the south of the country.