Rome, which had lost its central role in the administration of the empire, was sacked in 410 by the Visigoths led by Alaric I, but very little physical damage was done, most of which were repaired. What could not be so easily replaced were portable items such as artwork in precious metals and items for domestic use (loot). The popes embellished the city with large basilicas, such as Santa Maria Maggiore (with the collaboration of the emperors). The population of the city had fallen from 800,000 to 450–500,000 by the time the city was sacked in 455 by Genseric, king of the Vandals. The weak emperors of the fifth century could not stop the decay, leading to the deposition of Romulus Augustus on 22 August 476, which marked the end of the Western Roman Empire and, for many historians, the beginning of the Middle Ages. The decline of the city's population was caused by the loss of grain shipments from North Africa, from 440 onward, and the unwillingness of the senatorial class to maintain donations to support a population that was too large for the resources available. Even so, strenuous efforts were made to maintain the monumental centre, the palatine, and the largest baths, which continued to function until the Gothic siege of 537. The large baths of Constantine on the Quirinale were even repaired in 443, and the extent of the damage exaggerated and dramatised. However, the city gave an appearance overall of shabbiness and decay because of the large abandoned areas due to population decline. The population declined to 500,000 by 452 and 100,000 by 500 AD (perhaps larger, though no certain figure can be known). After the Gothic siege of 537, the population dropped to 30,000 but had risen to 90,000 by the papacy of Gregory the Great. The population decline coincided with the general collapse of urban life in the West in the fifth and sixth centuries, with few exceptions. Subsidized state grain distributions to the poorer members of society continued right through the sixth century and probably prevented the population from falling further. The figure of 450,000–500,000 is based on the amount of pork, 3,629,000 lbs. distributed to poorer Romans during five winter months at the rate of five Roman lbs per person per month, enough for 145,000 persons or 1/4 or 1/3 of the total population. Grain distribution to 80,000 ticket holders at the same time suggests 400,000 (Augustus set the number at 200,000 or one-fifth of the population).