Later, following the invitation of Grand Duke Gediminas, merchants and craftsmen began to move to Vilnius from the cities of the German Hanseatic League, France, Italy and Spain, and replaced the Lithuanian surnames with German, Polish, and Russian surnames. In the late 14th century, during the reign of Grand Duke Algirdas, Vilnius already had a Ruthenian quarter () in the present-day Latako and Rusų Streets, as the trade relations between the Grand Duchy of Lithuania and the Ruthenian principalities were quite well developed, therefore quite a few Ruthenian merchants lived there and the Ruthenian nobles had their residences in the quarter. The variety of nations in Vilnius was further increased by Grand Duke Vytautas the Great, who introduced Litvak Jews, Tatars and Crimean Karaites. After a few hundred years, the number of locals in Vilnius was smaller than the number of newcomers. However, according to an analysis of the tax registers of 1572, Lithuania proper had 850,000 residents, of which 680,000 (80%) were ethnically Lithuanians.