Modern Angola was populated predominantly by nomadic Khoi and San prior to the first Bantu migrations. The Khoi and San peoples were neither pastoralists nor cultivators, but rather hunter-gatherers. They were displaced by Bantu peoples arriving from the north in the first millennium BC, most of whom likely originated in what is today northwestern Nigeria and southern Niger. Bantu speakers introduced the cultivation of bananas and taro, as well as large cattle herds, to Angola's central highlands and the Luanda plain.