Phrygian rule was succeeded first by Lydian and later by Persian rule, though the strongly Phrygian character of the peasantry remained, as evidenced by the gravestones of the much later Roman period. Persian sovereignty lasted until the Persians' defeat at the hands of Alexander the Great who conquered the city in 333 BC. Alexander came from Gordion to Ankara and stayed in the city for a short period. After his death at Babylon in 323 BC and the subsequent division of his empire among his generals, Ankara, and its environs fell into the share of Antigonus.