After the fall of the Western Roman Empire in 476 AD, Rome was first under the control of Odoacer and then became part of the Ostrogothic Kingdom before returning to East Roman control after the Gothic War, which devastated the city in 546 and 550. Its population declined from more than a million in 210 AD to 500,000 in 273 to 35,000 after the Gothic War (535–554), reducing the sprawling city to groups of inhabited buildings interspersed among large areas of ruins, vegetation, vineyards and market gardens. It is generally thought the population of the city until 300 AD was 1 million (estimates range from 2 million to 750,000) declining to 750–800,000 in 400 AD, 450–500,000 in 450 AD and down to 80–100,000 in 500 AD (though it may have been twice this).