A 2014 study found that, based on ancestry informative markers (AIM), autosomal genetic ancestry in Cuba is 72% European, 20% African, and 8% Indigenous. Around 35% of maternal lineages derive from Cuban Indigenous People, compared to 39% from Africa and 26% from Europe, but male lineages were European (82%) and African (18%), indicating a historical bias towards mating between foreign men and native women rather than the inverse.