Southeast Asian features – specifically from the southern part of Borneo – are most predominant among the Merina of the central highlands, who form the largest Malagasy ethnic subgroup at approximately 26 percent of the population, while certain communities among the coastal peoples (collectively called ''côtiers'') have relatively stronger East African features. The largest coastal ethnic subgroups are the Betsimisaraka (14.9 percent) and the Tsimihety and Sakalava (6 percent each).