The remaining 5% of Nicaraguans are indigenous, the descendants of the country's original inhabitants. Nicaragua's pre-Columbian population consisted of many indigenous groups. In the western region, the Nahuas (Nicarao people) were present along with other groups such as the Chorotega people and the Subtiabas (also known as Maribios or Hokan Xiu). The central region and the Caribbean coast of Nicaragua were inhabited by indigenous peoples who were Macro-Chibchan language groups that had migrated to and from South America in ancient times, primarily what is now Colombia and Venezuela. These groups include the present-day Matagalpas, Miskitos, Ramas, as well as Mayangnas and Ulwas who are also known as Sumos. In the 19th century, there was a substantial indigenous minority, but this group was largely assimilated culturally into the mestizo majority. The Garifuna are also present, mainly on the Caribbean Coast. They are a people of mixed African and Indigenous descent.