Ecuador's mainstream culture is defined by its ''mestizo'' majority, and, like their ancestry, it is traditionally of Spanish heritage, influenced in different degrees by Amerindian traditions and in some cases by African elements. The first and most substantial wave of modern immigration to Ecuador consisted of Spanish colonists, following the arrival of Europeans in 1499. A lower number of other Europeans and North Americans migrated to the country in the late 19th and early 20th centuries and, in smaller numbers, Poles, Lithuanians, English, Irish, and Croats during and after the Second World War.