bombella apis bombella apis bombella apis is a gram-negative, strictly aerobic, non-spore-forming, rod-shaped and non-motile bacterium from the genus of bombella which has been isolated from the midgut of a bee (apis mellifera). bombella apis scientific classification domain: bacteria phylum: pseudomonadota class: alphaproteobacteria order: rhodospirillales family: acetobacteraceae genus: bombella species: b. apis binomial name bombella apisyun et al. 2017 type strain jcm 31623, kctc 52452, strain mrm1