plasmodium papernai plasmodium papernai plasmodium papernai is a parasite of the genus plasmodium subgenus novyella. as in all plasmodium species, p. papernai has both vertebrate and insect hosts. the vertebrate hosts for this parasite are birds. plasmodium papernai scientific classification domain: eukaryota clade: diaphoretickes clade: tsar clade: sar clade: alveolata phylum: apicomplexa class: aconoidasida order: haemospororida family: plasmodiidae genus: plasmodium species: p. papernai binomial name plasmodium papernaigrès and landau, 1997 taxonomy the parasite was first described by grès and landau in 1997.