occur on the glans of male and female spotted hyenas. in male dogs, the glans penis consists of two parts called the bulbus glandis and pars longa glandis. the glans of a fossa's penis extends about halfway down the shaft and is spiny except at the tip. in comparison, the glans of felids is short and spiny, while that of viverrids is smooth and long. the shape of the glans varies among different marsupial species. in most marsupials, the glans is divided, but male macropods have an undivided glans penis. the glans penis is also divided into two parts in platypuses and echidnas. the glans penis of the marsh rice rat is long and robust, averaging 7.3 mm (0.29 in) long and 4.6 mm (0.18 in) broad. winkelmann's mouse can most readily be distinguished from its close relatives by its partially corrugated glans penis. in thomasomys ucucha the glans penis is rounded, short, and small and is superficially divided into left and right halves by a trough at the top and a ridge at the bottom. most of the glans is covered with spines, except for an area near the tip. when erect, the glans of a horse's penis increases by 3 to 4 times. the urethra opens within the urethral fossa, a small pouch at the distal end of the glans. unlike the human glans, the glans of a horse's penis extends backwards on its shaft. males of racey's pipistrelle bat have a narrow, egg-shaped glans penis. the glans penis of a male cape ground squirrel is large with a prominent baculum.