hymenobacter terrenus hymenobacter terrenus hymenobacter terrenus is a gram-negative, non-spore-forming, short rod-shaped and non-motile bacterium from the genus of hymenobacter. it has been isolated from biological soil crusts from liangcheng, inner mongolia. hymenobacter terrenus scientific classification domain: bacteria phylum: bacteroidota class: cytophagia order: cytophagales family: hymenobacteraceae genus: hymenobacter species: h. terrenus binomial name hymenobacter terrenustang et al. 2015