dyadobacter crusticola dyadobacter crusticola dyadobacter crusticola is a gram-negative, psychrotolerant, aerobic and non-motile bacterium from the genus of dyadobacter which has been isolated from biological soil crusts from the colorado plateau in the united states. dyadobacter crusticola scientific classification domain: bacteria phylum: bacteroidota class: cytophagia order: cytophagales family: spirosomaceae genus: dyadobacter species: d. crusticola binomial name dyadobacter crusticolareddy and garcia-pichel 2005 type strain atcc baa-1036, cip 109009, cp183-8, dsm 16708, vtt e-072669