pseudomonas tuticorinensis pseudomonas tuticorinensis pseudomonas tuticorinensis is a gram-negative, marine denitrifying bacterium. the type strain is atcc 12230. pseudomonas tuticorinensis scientific classification domain: bacteria phylum: pseudomonadota class: gammaproteobacteria order: pseudomonadales family: pseudomonadaceae genus: pseudomonas species: p. tuticorinensis binomial name pseudomonas tuticorinensissreenivasan 1956