pontibacter rugosus pontibacter rugosus pontibacter rugosus is a gram-negative, rod-shaped, aerobic and motile bacterium from the genus of pontibacter which has been isolated from seawater from the gwangyang bay in korea. pontibacter rugosus scientific classification domain: bacteria phylum: bacteroidota class: cytophagia order: cytophagales family: hymenobacteraceae genus: pontibacter species: p. rugosus binomial name pontibacter rugosuskim et al. 2016 type strain jcm 31319, kctc 18739, kyw1030