comamonas granuli comamonas granuli comamonas granuli is a gram-negative, catalase-positive, oxidase-positive, non-spore-forming, motile, rod-shaped bacterium from the genus comamonas and family comamonadaceae, which was isolated from microbial granules. comamonas granuli scientific classification domain: bacteria phylum: pseudomonadota class: betaproteobacteria order: burkholderiales family: comamonadaceae genus: comamonas species: c. granuli binomial name comamonas granulikim et al. 2011 type strain kctc 12199, ko03, nbrc 101663