phyllobacterium catacumbae phyllobacterium catacumbae phyllobacterium catacumbae is a gram-negative, aerobic, motile bacteria with a polar tuft of flagella in the genus phyllobacterium. it was isolated from the tuff (a type of rock) walls of saint callixtus' catacomb in rome, italy. phyllobacterium catacumbae scientific classification domain: bacteria phylum: pseudomonadota class: alphaproteobacteria order: hyphomicrobiales family: phyllobacteriaceae genus: phyllobacterium species: p. catacumbae binomial name phyllobacterium catacumbaejurado et al. 2005 type strain cect 5680, cip 108923, csc19, lmg 22520 synonyms phyllobacterium callistus