peptide backbone, and the observation of congeners (pyoverdines from a single strain differing only in their ketoacid portions), nomenclature of pyoverdines remains rather tenuous and no single system has garnered universal acceptance. history 1850s: sédillot notes a blue-green discharge from surgical wound dressings. 1860: pyoverdine (although not so named) was extracted from wound dressings by fordos. 1862: lucke associates pyoverdine with bacilli observed under microscope. 1882: pseudomonas aeruginosa grown for first time in pure culture by carle gessard, reported in "on the blue and green coloration of bandages". gessard names the organism bacillus aeruginosa, after "aerugo", the latin word for verdigris. 1889: bouchard observes that injection of a rabbit infected with bacillus anthracis (causative agent of anthrax) with p. aeruginosa prevents formation of anthrax. 1889: bouchard discovers that pyoverdine fluoresces under ultraviolet light. 1948, 1952: first observations that concentrations of iron and pyoverdine are reciprocal. 1978: meyer and colleagues make first demonstration of role for pyoverdine in iron acquisition. 1980s–1990s: first structures and regulation of proverdine worked out 1999: first determination that pyoverdine fluorescence is quenched by iron binding. other uses pseudoverdine a compound related to pyoverdine, called pseudoverdine (formally known as 3-formylamino-6,7-dihydroxycoumarin) is also produced by some fluorescent pseudomonads. it is thought that pseudoverdine and pyoverdine may arise from a common precursor, 2,4,5-trihydroxyphenylalanine, which may condense with l-2,4-diaminobutyric acid to initiate pyoverdine production. pseudoverdine is relatively similar to pyoverdine in its fluorescence and other spectroscopic properties, and its ability to chelate ferric iron, albeit at much lower affinity. unlike pyoverdine, it is incapable of transporting iron into cells, likely due to the absence of the peptide chain. another dissimilarity is that pseudoverdine does not appear to be regulated by the same processes as pyoverdine.