secundilactobacillus secundilactobacillus secundilactobacillus is a genus of lactic acid bacteria containing secondary fermenters separated from the primary fermenters of lactobacillus after the latter group depletes hexoses and disaccharides. these gram-positive, rod-shaped heterofermenters have been isolated from beer and apple cider, and they are generally incapable of reducing fructose to mannitol. secundilactobacillus scientific classification domain: bacteria phylum: bacillota class: bacilli order: lactobacillales family: lactobacillaceae genus: secundilactobacilluszheng et al. 2020 type species secundilactobacillus malefermentans(farrow et al. 1989) zheng et al. 2020 species see text. species secundilactobacillus cladogram secundilactobacillus secundilactobacillus mixtipabuli secundilactobacillus silagincola secundilactobacillus silagei secundilactobacillus pentosiphilus secundilactobacillus odoratitofui secundilactobacillus similis secundilactobacillus collinoides secundilactobacillus paracollinoides secundilactobacillus kimchicus secundilactobacillus malefermentans secundilactobacillus oryzae outgroup holzapfelia phylogenetic relationships of twelve species reclassified by zheng et al. 2020 using whole-genome sequencing, zheng et al. 2020 reclassified twelve lactobacillus species into the novel genera secundilactobacillus. the list of prokaryotic names with standing in nomenclature (lpsn) currently recognizes fifteen member species: secundilactobacillus angelensis zhang et al. 2022 secundilactobacillus collinoides (carr and davies 1972) zheng et al. 2020 secundilactobacillus folii phuengjayaem et al. 2021 secundilactobacillus hailunensis liu et al. 2021 secundilactobacillus kimchicus (liang et al. 2011) zheng et al. 2020 secundilactobacillus malefermentans (farrow et al. 1989) zheng et al. 2020 secundilactobacillus mixtipabuli (tohno et al. 2015) zheng et al. 2020 secundilactobacillus odoratitofui (chao et al. 2010) zheng et al. 2020 secundilactobacillus oryzae (tohno et al. 2013) zheng et al. 2020 secundilactobacillus paracollinoides (suzuki et al. 2004) zheng et al. 2020 secundilactobacillus pentosiphilus (tohno et al. 2017) zheng et al. 2020 secundilactobacillus silagei (tohno et al. 2013) zheng et al. 2020 secundilactobacillus silagincola (tohno et al. 2017) zheng et al. 2020 secundilactobacillus similis (kitahara et al. 2010) zheng et al. 2020 secundilactobacillus yichangensis liu et al. 2021 these species have genomes sizes ranging from 1.85 mbp for s. oryzae to 3.62 mbp for s. collinoides, and their gc-content ranges from 41.03-47%.