papiine betaherpesvirus 4 papiine betaherpesvirus 4 papiine betaherpesvirus 4 (pahv-4) is a species of virus in the genus cytomegalovirus, subfamily betaherpesvirinae, family herpesviridae, and order herpesvirales. papiine betaherpesvirus 4 virus classification (unranked): virus realm: duplodnaviria kingdom: heunggongvirae phylum: peploviricota class: herviviricetes order: herpesvirales family: herpesviridae genus: cytomegalovirus species: papiine betaherpesvirus 4