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Lebanon now has "the highest per capita concentration of refugees worldwide", said the head of the UN refugee agency.
"For Lebanon, a small nation beset by internal difficulties, the impact is staggering," he added.
About 9.5 million people, almost half of Syria's population, have fled their homes since the start of the conflict.
More than 2.5 million have fled the country, with large numbers being taken in by Turkey, Jordan, Iraq, Egypt and others.
However Lebanon is bearing the biggest burden of all - the number of Syrian refugees there now amount to a quarter of the local population.
By Jim MuirBBC News, Beirut
Everybody knows that the real number of Syrian refugees in Lebanon is already well past the million mark, but the fact that that many have now been officially registered is yet another grim milestone as the conflict grinds on.
Lebanon's the smallest and most vulnerable of Syria's neighbours, yet it's taken in by far the largest number of refugees - one for every four of the country's own people.
That's a huge strain - the foreign minister said last month that it threatened Lebanon's very existence, and it's growing all the time, with about 2,500 new refugees being registered every day.
The UN has only received 14% of the funding it's asked for. That means that relief has to be cut back and carefully targeted to the most needy.
The BBC's Jim Muir in Beirut says this is a huge strain for Lebanon, the smallest and most vulnerable of Syria's neighbours.
Last month the Lebanese foreign minister said the crisis was "threatening the existence" of his country.
"The Lebanese people have shown striking generosity, but are struggling to cope," Mr Guterres said.
The UN's refugee agency, UNHCR, notes that the influx of refugees is accelerating.
"Every day, UNHCR in Lebanon registers 2,500 new refugees: more than one person a minute," the agency said in a statement.
International aid agencies are finding it increasingly difficult to cope with the flow of refugees.
UNHCR has only received 14% of the $6.5bn (Â£4bn; 4.7bn euros) funding it has asked for.
The hardships many refugees are facing was dramatically highlighted last week when a mother with a sick husband and four children set fire to herself in protest at not receiving help. She was critically injured.
The civil war in Syria between Sunni-led rebels and the government of Bashar al-Assad has also fuelled sectarian tensions in Lebanon, which has a large Sunni and Shia Muslim population.
Hundreds of people in Beirut and other parts of the country have been killed in violence between opponents and supporters of President Assad.
The number of refugees who have fled Syria and registered in Lebanon has surpassed the "devastating milestone" of one million, the UN says.

Nigel Wilson, 42, from Bingham, near Nottingham, appeared at Westminster Magistrates' Court accused of 17 breaches of the Air Navigation Order.
It is the first prosecution of its kind in England to follow a police-led investigation.
Mr Wilson admitted two counts, but will go on trial for the remaining 15.
Mr Wilson, of Rockingham Grove, admitted two counts of not maintaining visual, unaided contact with an unmanned surveillance aircraft to monitor its flight path.
They relate to incidents over the north bank of the River Thames on September 27 and during Manchester City v Tottenham Hotspur at the Etihad Stadium, Manchester on October 18.
He denied other charges of flying a "small unmanned surveillance aircraft" without permission during football matches at various grounds.
They included Arsenal's Emirates Stadium, Liverpool's Anfield ground as well as Nottingham Forest's City Ground, Derby County's iPro stadium and the King Power Stadium in Leicester.
The charges also claim he breached regulations by failing to maintain direct visual contact with the drone as he flew it near landmarks including Buckingham Palace and the Palace of Westminster.
The majority of the offences are said to have happened in September last year.
Prosecutor Edward Aydin said: "These were extremely busy areas.
"The fear of the police and the Civil Aviation Authority is that it [the drone] is going to fall out of the sky and hit someone."
The Civil Aviation Authority successfully brought two similar cases to court in 2013 and 2014.
However, this is the first time someone has been prosecuted by the Crown Prosecution Service after a police operation.
Mr Wilson is due to appear at the same court on Friday so that a date for his trial can be set.
A man alleged to have illegally flown a camera drone over packed football stadiums and London tourist attractions has appeared in court.

He had been nominated last week for outstanding guest actor in a comedy for his role as Jeff Kane in the HBO show.
The category is open to actors who have appeared in less than 50% of a programme's current series.
MacNicol appeared in five of the 10 episodes of Veep's latest season - exactly 50%.
The Television Academy told The Hollywood Reporter: "Upon review of the category, we have determined that Peter MacNicol unfortunately exceeds the number of episodes permissible for inclusion."
The academy acknowledged MacNicol's entry had been eligible at the time of submission, but added he had subsequently appeared in a further Veep episode.
"Unfortunately, that additional appearance places him in 50% of the season's episodes and makes him no longer eligible to compete in the guest actor category," it said.
"This decision is in no way a diminishment of Mr MacNicol's stellar performance on this season of Veep."
The academy said a replacement nominee would be announced shortly.
Veep - a comedy produced for subscription network HBO - has just concluded its fifth season.
The show was conceived by The Thick of It creator Armando Iannucci as a political satire focusing on the fictional US vice president and her staff.
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Veep actor Peter MacNicol has had his Emmy nomination withdrawn after the Television Academy ruled his performance was ineligible.

