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Lebanon now has "the highest per capita concentration of refugees worldwide", said the head of the UN refugee agency.
"For Lebanon, a small nation beset by internal difficulties, the impact is staggering," he added.
About 9.5 million people, almost half of Syria's population, have fled their homes since the start of the conflict.
More than 2.5 million have fled the country, with large numbers being taken in by Turkey, Jordan, Iraq, Egypt and others.
However Lebanon is bearing the biggest burden of all - the number of Syrian refugees there now amount to a quarter of the local population.
By Jim MuirBBC News, Beirut
Everybody knows that the real number of Syrian refugees in Lebanon is already well past the million mark, but the fact that that many have now been officially registered is yet another grim milestone as the conflict grinds on.
Lebanon's the smallest and most vulnerable of Syria's neighbours, yet it's taken in by far the largest number of refugees - one for every four of the country's own people.
That's a huge strain - the foreign minister said last month that it threatened Lebanon's very existence, and it's growing all the time, with about 2,500 new refugees being registered every day.
The UN has only received 14% of the funding it's asked for. That means that relief has to be cut back and carefully targeted to the most needy.
The BBC's Jim Muir in Beirut says this is a huge strain for Lebanon, the smallest and most vulnerable of Syria's neighbours.
Last month the Lebanese foreign minister said the crisis was "threatening the existence" of his country.
"The Lebanese people have shown striking generosity, but are struggling to cope," Mr Guterres said.
The UN's refugee agency, UNHCR, notes that the influx of refugees is accelerating.
"Every day, UNHCR in Lebanon registers 2,500 new refugees: more than one person a minute," the agency said in a statement.
International aid agencies are finding it increasingly difficult to cope with the flow of refugees.
UNHCR has only received 14% of the $6.5bn (Â£4bn; 4.7bn euros) funding it has asked for.
The hardships many refugees are facing was dramatically highlighted last week when a mother with a sick husband and four children set fire to herself in protest at not receiving help. She was critically injured.
The civil war in Syria between Sunni-led rebels and the government of Bashar al-Assad has also fuelled sectarian tensions in Lebanon, which has a large Sunni and Shia Muslim population.
Hundreds of people in Beirut and other parts of the country have been killed in violence between opponents and supporters of President Assad.
The number of refugees who have fled Syria and registered in Lebanon has surpassed the "devastating milestone" of one million, the UN says.

Germany rejected a Greek request for a six-month extension to its loan programme without austerity reforms, calling it a "Trojan horse" proposal.
Japan's benchmark Nikkei 225 and broader Topix both closed 0.3% up at 18,321 and 1,499.28 respectively.
Trading in the rest of Asia was quiet over the Lunar New Year holiday.
Markets in China, Hong Kong, Malaysia, Singapore, South Korea, Taiwan and Vietnam remain closed.
In Sydney, the S&P/ASX 200 ended 0.4% lower at 5,881.50 because of a decline in mining and energy stocks. New Zealand stocks rose 0.4% to finish at 5,748.95.
Andrew Kenningham, senior global economist at Capital Economics said a Greek exit from the eurozone would be felt primarily through the financial markets.
"The shock of 'Grexit' finally taking place would be likely to dampen global risk appetite, leading to a sell-off in global equities and some emerging market assets," he wrote in a report.
"On the other hand, there would be an increase in demand for safe havens, including non-euro government bonds and gold."
Gold prices rose by more than 1% and the metal was trading at about $1,208.70 an ounce in the US spot market overnight.
Japanese stocks rallied further on Friday, despite concerns Greece may be forced to leave the eurozone should it fail to negotiate a new bailout deal.

Eddie Todd, an independent in Doncaster's mayoral race, was given a ticket for having unauthorised blue lights on his vehicle which he denies.
Film of the incident in Doncaster on Wednesday has been shared on social media and Mr Todd claims it may have jeopardised his campaign.
South Yorkshire Police said it did not comment on individual cases.
Read more about this and other stories across Yorkshire
Mr Todd's car is used as a private ambulance and in the video a passer-by asks an officer why he was being issued with a ticket.
The officer said he was not allowed to use blue lights as the car was a private vehicle and he was not an ambulance driver.
Mr Todd told the BBC: "I was set up and someone took a video."
He said he would consider taking the incident "further" with the police and was planning to make a formal complaint.
Mr Todd came fourth in the mayoral race and received 5,344 votes (8.3%). Labour's Ros Jones was elected with 32,631 votes (50.9%).

A mayoral candidate has claimed he was "set up" after he was filmed being given a fixed penalty notice.

