thiophaeococcus fuscus thiophaeococcus fuscus thiophaeococcus fuscus is a gram-negative and motile bacterium from the genus of thiophaeococcus has been isolated from black sand from a lagoon. thiophaeococcus fuscus scientific classification domain: bacteria phylum: pseudomonadota class: gammaproteobacteria order: chromatiales family: chromatiaceae genus: thiophaeococcus species: t. fuscus binomial name thiophaeococcus fuscusdivyasree et al. 2014 type strain kctc 15337, nbrc 109958, strain ja633