chlorobaculum tepidum chlorobaculum tepidum chlorobaculum tepidum, previously known as chlorobium tepidum, is an anaerobic, thermophilic green sulfur bacteria first isolated from new zealand. its cells are gram-negative and non-motile rods of variable length. they contain chlorosomes and bacteriochlorophyll a and c. chlorobaculum tepidum scientific classification domain: bacteria phylum: chlorobiota class: "chlorobia" order: chlorobiales family: chlorobiaceae genus: chlorobaculum species: c. tepidum binomial name chlorobaculum tepidum(wahlund et al. 1996) imhoff 2003 synonyms chlorobium tepidum wahlund et al. 1996 natural habitat and environmental requirements like other green sulfur bacteria c. tepidum requires light and specific compounds to perform anoxygenic photosynthesis. the compounds that c. tepidum requires reduced sulfur compounds (h2s, s0, thiosulfate), hydrogen gas (h2,) or ferrous iron (fe2+). to fulfill their metabolic requirements, they reside primarily in anaerobic sulfur rich environments such as anaerobic levels of stratified lakes and lagoons, anaerobic levels of layered organic bacterial mats, and in hot springs where there is abundant sulfur. c. tepidum and other green sulfur bacteria also play a large role within the carbon and sulfur cycles. within the sulfur cycle, they contribute to the oxidative branch by oxidizing reduced sulfur compounds. within anaerobic sediment layers c. tepidum is able to couple carbon and sulfur cycling in a metabolically favorable way. photosynthetic mechanism as it was mentioned before, c. tepidum performs anoxygenic photosynthesis. within each cell there are 200-250 chlorosomes that are attached to the cytoplasmic side of reaction centers inserted within the inner cell membrane. the ellipsoidal shaped complexes act as light harvesting antenna to capture energy. within each chlorosome are 215,000 ± 80,000 bacteriochlorophyll c that act as pigment molecules and absorb unique wavelengths of light relative to their color. light energy is harvested by the chlorosomes and used in conjunction with h2, reduced sulfur compounds, or ferrous iron to preform redox reactions and provide energy to fix co2 via the reverse tricarboxcylic acid cycle. genome structure c. tepidum contains a genome that contains 2.15 mbp, within there are a total of 2,337 genes (of these genes, there are 2,245 protein coding genes and 56 trna and rrna coding genes). it's synthesis of chlorophyll a and bacteriochlorophylls a and c make it a model organism used to elucidate the biosynthesis of bacteriochlorophylls c. several of its carotenoid metabolic pathways (including a novel lycopene cyclase) have similar counterparts in cyanobacteria.