holophagaceae holophagaceae the class holophagaceae is a family of strictly anaerobic gram negative marine bacteria in the phylum acidobacteriota. holophagaceae scientific classification domain: bacteria phylum: acidobacteriota class: holophagae order: holophagalesfukunaga et al. 2008 family: holophagaceaefukunaga et al. 2008 genera geothrix coates et al. 1999 holophaga liesack et al. 1995