gramella echinicola gramella echinicola gramella echinicola is a gram-negative and halophilic bacterium from the genus of gramella which has been isolated from the sea urchin strongylocentrotus intermedius from the sea of japan. gramella echinicola scientific classification domain: bacteria phylum: bacteroidota class: flavobacteriia order: flavobacteriales family: flavobacteriaceae genus: gramella species: g. echinicola binomial name gramella echinicolanedashkovskaya et al. 2005 type strain kmm 6050