crinalium epipsammum crinalium epipsammum crinalium epipsammum is a filamentous, non-heterocystous, terrestrial cyanobacterium. the species is highly drought-resistant and was first identified from its role in forming crusts on coastal sand dunes in the netherlands. crinalium epipsammum scientific classification domain: bacteria phylum: cyanobacteria class: cyanophyceae order: oscillatoriales family: gomontiellaceae genus: crinalium species: c. epipsammum binomial name crinalium epipsammumwinder, stal & mur, 1992