pusillimonas harenae pusillimonas harenae pusillimonas harenae is a gram-negative, oxidase- and catalase-positive, motile bacterium of the genus pusillimonas, isolated from beach sand on the taean coast in south korea. pusillimonas harenae scientific classification domain: bacteria phylum: pseudomonadota class: betaproteobacteria order: burkholderiales family: alcaligenaceae genus: pusillimonas species: p. harenae binomial name pusillimonas harenaepark et al. 2011 type strain b201, jcm 16917, kacc 14927