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It is the center of body metabolism and contains the vast bulk of digestive , reproductive , circulatory , and excretory organs . The Cameroon study was conducted in the dry season , while the Bioko study was conducted in the wet season , so the studies are not completely comparable , but the richness of the Bioko fauna is undeniable . This family of two-clawed running spiders has recently been separated from the Clubionidae . The skull is the most complex segment of the mammal skeleton .