[word=nuclei] [lemma=□]
[word=nuclei] [lemma=which]
However , a point which Davis mentions but does not emphasize ( in contrast to Le Doux 1993 ; and Helmstetter 1992 ) is that the amygdala is clearly involved in unconditioned fear as well as conditioned fear and is one of the higher levels of the system of nuclei which becomes generally active in a wide variety of aversive situations ( e.g . However , the thalamic nuclei which provide input to the posterior cingulate include ones which project to the frontal cortex ; and , at least with input from the pain pathways , the anterior cingulate acts as the primary sensory cortex ( Rainville et al. 1997 ) . Another alternative view is that the subcortical outputs of the amygdala activate target nuclei which not only control specific responses via descending connections but also ( in the cases of the basal forebrain , substantia nigra , lateral dorsal tegmental nucleus , and pedunculopontine tegmental nucleus ) control a variety of different types of attention via ascending connections to the diencephalon and telencephalon ( Gallagher and Holland 1994 ; Gallagher and Chiba 1996 ) . The cells are large and polygonal with abundant pale eosinophilic granular cytoplasm and small vesicular nuclei which are centrally located within the cell .