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For a discussion of the roles of microtubule-associated proteins and motor proteins in the generation of organized microtubule arrays , we direct the reader to excellent recent reviews2 , 3.The centrosome — a central organizerSince its description by Theodor Boveri more than 100 years ago , the centrosome has attracted special attention because of its prominent location at the centre of the microtubule cytoskeleton and because it contains one of the most recognizable structures found in eukaryotic cells , the centriole .