The promoter region between -173 and -60 contained the strongest activating elements .
the promoter region between -173 and -60 contained the strongest activating elements .
O B-DNA_domain_or_region I-DNA_domain_or_region O B-DNA_domain_or_region I-DNA_domain_or_region I-DNA_domain_or_region O O O B-DNA_domain_or_region I-DNA_domain_or_region O