Both Sp1 and Sp3 bound to the 5' portion of this region , whereas a PMA -inducible factor ( PIF ) mainly bound to its 3' portion and bound to the Sp binding motifs as well .
both sp1 and sp3 bound to the 5' portion of this region , whereas a pma -inducible factor ( pif ) mainly bound to its 3' portion and bound to the sp binding motifs as well .
O B-protein_molecule O B-protein_molecule O O O B-DNA_domain_or_region I-DNA_domain_or_region O O O O O O B-other_organic_compound O O O B-protein_family_or_group O O O O O B-DNA_domain_or_region I-DNA_domain_or_region O O O O B-DNA_family_or_group I-DNA_family_or_group I-DNA_family_or_group O O O