Author contributions
D.L., N.H.
J.T., J.H.C., S.X.C., K.P., M.G., J.H., K.B., J.B., T.O., M.H., G.S., A.J.G., A.S., S.D., R.L., C.D.C., O.Y., G.A.G., A.R.P., N.H., and R.B.C. participated in the design, execution and/or interpretation of the reported experiments or results. J.T., J.H.C., S.X.C., K.P., V.J., P.S., A.M., D.L., M.S., J.N.A., T.A.A., J.W.C., A.C.E., P.C.E., S.J.K., N.J.M., J.A.M., D.P.R., M.B.Y., K.K., E.E.V.S., I.B., G.C., J.J., W.B.B., E.W., A.G.M. and I.J.F. participated in the sample acquisition or data analysis. J.T. and R.B.C. wrote the paper, with all authors contributing to writing and providing feedback. N.H. and R.B.C. supervised all aspects of the research.