Author contributions
K.P.W., N.R., and C.M.D. designed the experiments; C.M.D., S.J.M, and K.P.W. collected data; all authors analyzed data and edited the paper; M.G., C.M.D., and K.P.W. wrote the manuscript; K.P.W., N.R., and C.M.D. obtained funding and supervised the research.