Author contributions
JJ, CML
CML
JJ and SHK designed the study. JJ, JHP, and HY collected and analyzed the data. JJ wrote the main manuscript text and JHP prepared figure. YJL, EOK, CML, MNK, MWJ, SCY, and SHK reviewed and revised the manuscript. This study was presented in part in ASM Microbe, 2022, Jun 12, Washington DC (Abstract number CIV-2578). All authors read and approved the final manuscript.