Author contributions
LIU
LI Qingkai and LIU Ying performed the experiments and wrote the article; WANG Yanan, SHAN Xin and LIU Chunxiao collected and analysed data; CAO Jinglin and LI Zhihua performed the experiments and collected and analysed data; XU Guanjie and DOU Jian designed the study and reviewed the article; WANG Qiujun revised the article; QIE Xiaojuan conceived, reviewed and funded the article. All authors read and approved the final manuscript as submitted.