Author contributions
AML
AML
PL, JPV and GI participated in the conception and design of the study. GI, TM, GM and BM performed the fieldwork and wrote the first draft manuscript. AML, VM, JRM, SL participated in the development of the computer software, the analysis, interpretation of data and revision of manuscript. JRM, NMM and AML analysed, interpreted the data and revised the manuscript. FTM, JPVand PL participated in the revision of manuscript. All authors read and approved the final manuscript.