Author contributions
MM
JC and CWD designed the study, processed and analyzed phenotype and genotype data, performed statistical analyses, and wrote the manuscript. KS and RMH analyzed data and wrote the manuscript. E Place, EZ, and KMB processed genotype data and analyzed variants. MM processed phenotype and genotype data. YZ performed statistical analyses. E Pierce designed the study, analyzed phenotype and genotype data, wrote the manuscript, and obtained funding. Co–first author order was determined by the authors’ relative contributions to the content, analysis, and writing of the manuscript.