Author contributions
V.E.R.P. and D.R. designed the research. V.B.S.-H., P.S., B.H., E.P., K.H.M.R., E.E.-T., M.d.L., K.S., D.J.W. and F.S. performed the experiments. V.E.R.P., D.R., Y.-T.C., P.S., J.M., K.S., E.J., L. Johansson and D.J.W. analysed the data. V.E.R.P., P.A., M.d.L., L.H., L. Jermutus and R.E. supervised the project. All authors contributed to writing the manuscript and approved the manuscript for submission.