Author contributions
HLH, CJD, RDR
CJD
MC, GIW, RRF, SCS, and MJB designed research; MC, JKF, RDR, ASO, LKD, and MJB performed research; JS, CJD, SJH, PP, LG, HLH, and RDT contributed resources; MC, JKF, KLO, ASO, MG, LS, KHW, and MJB analyzed data; MC, SCS, and MJB wrote paper; and all authors approved the final version of the manuscript.