Author Contributions
J.-E.H. set up the experiments, performed the data analysis, and drafted the manuscript as the first author. J.-Y.P. and H.J. administrated the project and reviewed the manuscript. M.H.J., K.E. and H.S.M. participated in the project and conducted the experiments. Y.J.K. designed the study, performed the data analysis, and drafted the manuscript as the corresponding author. All authors have read and agreed to the published version of the manuscript.