Author contributions
There are 3 first authors in this manuscript and they have equally contributed to this project. T.Z., G.Y., and C.L. were responsible for the design of the project and all data sorting, and writing articles. W.W., C.T., X.C., Q.H., Z.C., P.S., Y.D., H.X., and L.G. were responsible for data analysis and writing articles. D.C., Y.S., Q.C., and W.D. were responsible for sample sequencing and data analysis. Z.Y. supervised the work. All authors read and approved the final manuscript.