Author contributions
DRD, JJ, JP
JJ and JP conceived and designed research; JJ, JDS, DRD, KM, JRN, and JP performed experiments; JJ and JP analyzed data; JJ, JDS, and JP interpreted results of experiments; JJ and JP prepared figures; JJ, JDS, and JP drafted the manuscript; JJ, JDS, DRD, KM, JRN, and JP edited and revised the manuscript; and JJ, JDS, DRD, KM, JRN, and JP approved final version of the manuscript.