Author contributions
SCA, MPD
MOA conceived the study. MOA, BG, JLAN and DS developed the study protocol. MOA, JLAN and DS supervised the data collection. IMM, EBF, IAM coordinated the fieldwork. AS, FBS and MAL managed the datasets. ACL, MPD, NAG, AD performed the laboratory analysis. SCA analysed the data. MOA developed the first draft of the manuscript. BG, JLAN and DS reviewed the manuscript critically for important intellectual contents. All authors reviewed and approved the final draft of the manuscript.