Contributors
NP, FWM, AM, MR, JFM, and NSM conceived and designed the study. NSM led and performed the field work with support from JM, MJ, and supervision from NP. NSM and MAK conducted and analysed the data. EM and EL performed the data curation. NSM, OM, BS, and RK performed the molecular analysis with support from LAM. NP, JFM, AM, MR, LAM, MAK, and FWM supported data interpretation. NSM wrote the original paper. NP, MAK, and LAM critically reviewed the manuscript. All authors read and approved the final manuscript. NSM, MK, and NP had accessed and verified the data and all authors had full access to all the data in the study