Author contributions
SEB, AMV, JW, NS, PWF, TDS, GH and JC designed research. SEB, JW, GH and TDS conducted research. AM, KMB and FA performed statistical analysis. KMB, SEB, AMV, JW, TDS, NS, FA, ERL and LMD wrote the paper. SEB, TDS and KMB had primary responsibility for final content. All authors read and approved the final manuscript.