Author contributions
J.Y. and K.H.H. designed the study. H.W.K. and J.L. wrote the manuscript. H.W.K. and S.-J.H. analyzed the data. J.Y. reviewed and edited the manuscript. H.W.K., J.L., B.S.K., K.H.H., and J.Y. analyzed the data and contributed to the discussion. J.Y. is the guarantor of this work and takes responsibility for the integrity of the data. All authors critically revised the manuscript for key intellectual content and approved the final version.