He graduated from the University of Tartu in 1975. His earlier work dealt with ethology and field ecology. He has studied the mechanisms of species coexistence in species-rich communities and developed mathematical modelling in ecophysiology.
A scholar from the University of Tartu graduated in 1975. Initially, his work centered on animal behavior (ethology) and outdoor research (field ecology). He examined how different species live together in complex environments and created mathematical frameworks to study physiological responses (ecophysiology).