Though often referred to as a cross between the lemon and citron, a recent genomic analysis showed it to solely contain pomelo (Citrus maxima) and citron (Citrus medica) DNA, perhaps being an F2 hybrid.
The genetic analysis suggests that this citrus fruit is not a cross between lemon and citron but rather a result of hybridization between pomelo (Citrus maxima) and citron (Citrus medica). This finding could be explained by the presence of F2 hybrid DNA in its genome.