Species of Geocrinia develop initially in the egg then hatch when sufficient rain falls for them to complete their larval development in water, while Anstisia go through the tadpole stage entirely on land in the egg capsule.
The passage describes two different amphibian species with unique developmental patterns: Geocrinia which develop in eggs that hatch into larvae living in water, and Anstisia which have a tadpole stage entirely on land within the egg capsule.