The papyrological hyphen () showed a group of letters should be read together as a single word, and the hypodiastole showed that they should be taken separately. Compare "" ("whatever") to "" ("...that...").
The use of a hyphen in ancient Egyptian writing (papyrological hyphen) meant that a sequence of letters was read as one unit, whereas the hypodiastole indicated that these letters should be treated individually.  This is compared to the difference between "whatever" and "..." ("...that...").