They later applied the same testing to complex cells, but used the Y cell (subunit) model instead. This model stated that each subunits could respond differently, but the converted responses would be offset in time, so it would sum to a constant value.
They later applied the same testing to complex cells, but employed the Y cell (subunit) model instead. This model asserted that each subunit could respond differently, yet the converted responses would be offset in time, resulting in a constant value summation.