A second biotype (L-biotype) was  found on liana vines in Ecuador; subsequently the host was identified as Arrabidaea verrucosa (Bignoniaceae) but witches' broom symptoms have not been observed on this host.
A secondary strain (L-biotype) was discovered growing on liana vines in Ecuador. Following the identification of its host plant as Arrabidaea verrucosa (Bignoniaceae), it became clear that witches' broom symptoms are not present on this particular host.