This can be quantified using relative failure force, which is the bone failure force of a vertebra divided by the body weight of the pterosaur that it belongs to, estimated at  for Arambourgiania and Hatzegopteryx.
This can be measured using relative failure force, which is the bone strength of a vertebra divided by the body weight of the pterosaur it belongs to, estimated at  for Arambourgiania and Hatzegopteryx.