AbstractBased on recent achievements in phylogenetic studies of theBrassicaceae, a novel infrafamilial classification is proposed that includes major improvements at the subfamilial and supertribal levels. Herein, the family is subdivided into two subfamilies,Aethionemoideae(subfam. nov.) andBrassicoideae. TheBrassicoideae, with 57 of the 58 tribes ofBrassicaceae, are further partitioned into five supertribes, including the previously recognizedBrassicodaeand the newly establishedArabodae,Camelinodae,Heliophilodae, andHesperodae. Additional tribus-level contributions include descriptions of the newly recognizedArabidopsideae,Asperuginoideae,Hemilophieae,Schrenkielleae, and resurrection of theChamireaeandSubularieae. Further detailed comments on 17 tribes in need of clarifications are provided.