Explorer-naturalists such as Alexander von Humboldt investigated the interaction between organisms and their environment, and the ways in which this relationship depends on geography, laying the foundations for biogeography, ecology, and ethology.
Ndị nnyocha na ndị agụmaọnatarachi dịka Alexander von Humbodlt lebara ányá n'ime mmekọrịta dị n'etiti ndịorganism na gburugburu ha, n' ụzọ mmekọrịta a dabere n'esereseanị, nke debere ntọala maka esereseanịndụ, agụmagburugburu n'agụmàgwànụmanụ.