Meadows in the high end of the salinity gradient tend to be more productive (Bostrom et al., Key species of the macrozoobenthic community in the Baltic proper include Macoma balthica, Halicryptus spinulosus, Marenzelleria arctia and Saduria entomon, whereas in the Bothnian Sea, cold-water dominating species include Monoporeia affinis, Pontoporeiafemorata and Saduria entomon. Multiple pressures from agricultural landuse and maritime traffic (HELCOM, 2009) has resulted in large environmental changes during the last 100 years.