Digital literacies have fast become indispensable for productive engagement, competency, and citizenship in a rapidly changing world. After-school programs represent an important venue where many young people can develop a mastery of digital literacies, encompassing both the creative and responsible use of a broad range of new media. This paper discusses relational habitus (the configuration of self, tools, tasks, and others in a specific activity) in the development of digital literacies among youth in a network of after-school programs called University-Community Links (UC Links). A collaborative effort among university and local community partners throughout California, UC Links provides informal learning activities that enable underserved K-12 youth to develop their digital literacies. After presenting UC Links' approach to informal learning after school, we offer a cognitive ethnography, describing how a distinctive relational habitus configures the links among self, tools, tasks, and others in infor...