Context: The higher-level phylogeny of the arthropods continues to be a matter of debate and research. In 2008, researchers at Tufts University uncovered what they believe is the world's oldest known full-body impression of a primitive flying insect, a 300 million-year-old specimen from the Carboniferous period. The oldest definitive insect fossil is the Devonian Rhyniognatha hirsti, from the 396-million-year-old Rhynie chert. It may have superficially resembled a modern-day silverfish insect. This species already possessed dicondylic mandibles (two articulations in the mandible), a feature associated with winged insects, suggesting that wings may already have evolved at this time. Thus, the first insects probably appeared earlier, in the Silurian period.

Question: From what period do researchers believe that the oldest known full-body impression of a flying insect originates?
Answer: Carboniferous

Question: What is believed to be the oldest known insect fossil?
Answer: Devonian Rhyniognatha hirsti

Question: In what year did Tufts researchers reportedly uncover what is believed to be the oldest known impression of a flying insect?
Answer: 2008

Question: What is the term for a mandible with two articulations?
Answer: dicondylic

Question: With what type of insect is a dicondylic mandible associated?
Answer: winged

Question: What higher-level is constantly up for debate and research?
Answer: phylogeny

Question: The higher-level phylogeny is of the what?
Answer: arthropods

Question: What kind of impression has been uncovered by a University?
Answer: full-body

Question: A University discovered a primitive flying what?
Answer: insect

Question: How old is the primitive plying insect found by the University?
Answer: 300 million-year-old